2016 in paleontology

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Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2016.

Plants

Main article: 2016 in paleobotany

Cnidarians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Aphyllum pachystele[4]

Sp. nov

Valid

Munson & Jell

Silurian (late Wenlock)

Jack Formation

 Australia

A rugose coral belonging to the family Tryplasmatidae.

Aulokoninckophyllum potyi[5]

Sp. nov

Valid

Somerville, Rodríguez & Said

Carboniferous

Azrou-Khenifra Basin

 Morocco

An aulate coral.

Bjarmia[6]

Gen. et sp. nov

Valid

Grazhdankin

Late Ediacaran (~550 Ma)

Erga Formation

 Russia

A jellyfish-like organism, a possible relative of scyphomedusae. Genus includes new species B. cycloplerusa.

Calceola collini[7]

Sp. nov

Valid

Wright, Plusquellec & Gourvennec

Devonian (Eifelian)

Saint-Fiacre Formation

 France

An operculate coral belonging to the family Calceolidae; a species of Calceola.

Cambroctoconus coreaensis[8]

Sp. nov

Valid

Park et al.

Cambrian

Daegi Formation

 South Korea

A stem-cnidarian.

Caninophyllum charli[9]

Sp. nov

Valid

Denayer

Carboniferous (Tournaisian)

Yılanlı Formation

 Turkey

A rugose coral belonging to the family Bothrophyllidae; a species of Caninophyllum.

Dokophyllum hillae[4]

Sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Ketophyllidae.

Edaphophyllum zhaotongense[10]

Sp. nov

Valid

Liao & Ma

Devonian (Emsian)

 China

A rugose coral.

Eoaequorea[11]

Gen. et sp. nov

Valid

Tang et al.

Ediacaran

Doushantuo Formation

 China

A member of Hydrozoa of uncertain phylogenetic placement. The type species is Eoaequorea xingi.

Eosiderastrea[12]

Gen. et comb. et 2 sp. nov

Valid

Löser

Cretaceous (late Valanginian to Santonian)

 Austria
 France
 Germany
 Greece
 Iran
 Italy
 Japan
 Mexico
 Poland
 Spain
 Ukraine
 United States
 Uzbekistan

A stony coral belonging to the family Siderastreidae. The type species is "Siderastrea" cuyleri Wells (1932); genus also includes "Phyllocoenia" cotteaui Orbigny (1850), "Diploastrea" crassa Kuzmicheva (1980), "Diploastrea" crassicostata Morycowa & Masse (1998), "Phyllocoenia" cyclops Felix (1891), "Confusastrea" dollfusi Prever (1909), "Confusastrea" felixi Prever (1909); "Stephanocoenia" grandipora Orbigny (1849), "Diploastrea" harrisi Wells (1932), "Diploastrea" hilli Wells (1933), "Montastraea" nagaoi Eguchi (1951), "Pleurocora" reussi Milne Edwards (1857), "Diplocoenia" splendida Prever (1909), "Diploastrea" vaughani Wells (1933) and "Plesiofavia" villaltai Reig Oriol (1991), as well as new species E. paragrandipora Löser (2016) and E. stefani Löser (2016).[13]

Famastraea[14]

Gen. et sp. nov

Valid

Berkowski, Zapalski & Wrzołek

Devonian (Famennian)

 Poland

A coral belonging to the group Rugosa and the family Campophyllidae. The type species is Famastraea catenata.

Fomichevella najafi[15]

Sp. nov

Valid

Badpa et al.

Carboniferous (Bashkirian)

Sardar Formation

 Iran

A member of Rugosa belonging to the group Stauriida and the family Kleopatrinidae.

Gertholites haikawai[16]

Sp. nov

Valid

Shuji & Masayuki

Early Carboniferous

Akiyoshi Limestone Group

 Japan

A tabulate coral.

Gerviphyllum[7]

Gen. et comb. nov

Valid

Wright, Plusquellec & Gourvennec

Early Devonian (likely late Lochkovian to early Pragian)

 France

An operculate coral belonging to the family Calceolidae; a new genus for "Calceola" gervillei Bayle (1878).

Heintzella fluegeli[15]

Sp. nov

Valid

Badpa et al.

Carboniferous (Bashkirian)

Sardar Formation

 Iran

A member of Rugosa belonging to the group Stauriida and the family Kleopatrinidae.

Kullingia rotadiscopsis[11]

Sp. nov

Valid

Tang et al.

Ediacaran

Doushantuo Formation

 China

A member of Hydrozoa of uncertain phylogenetic placement; a species of Kullingia.

Madrepora mitsukurii[17]

Sp. nov

Valid

Shuji, Shigeyuki & Eiji

Miocene

Katsuta Group

 Japan

A madrepore.

Mesoconularia (?) lukesi[18]

Sp. nov

Valid

Mergl, Ferrova & Frýda

Devonian (late Emsian)

Suchomasty Limestone

 Czech Republic

A member of Conulariida, possibly a species of Mesoconularia.

?Michelinia vinni[19]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Micheliniidae; possibly a species of Michelinia.

Multicarinophyllum vepreculatum[4]

Sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Amsdenoididae.

Palaeosiderofungia[12]

Gen. et comb. nov

Valid

Löser

Cretaceous (Aptian to Santonian)

 Austria
 Czech Republic
 France
 Germany
 Greece
 Serbia

A stony coral belonging to the family Siderastreidae; a new genus for "Thamnasteria" exigua Reuss (1854).

Potyphyllum[20]

Gen. et comb. nov

Valid

Coen-Aubert

Devonian (late Frasnian)

 Belgium
 Czech Republic
 France
 Germany
 Poland
 Russia
 United Kingdom

A rugose coral belonging to the family Phillipsastreidae. The type species is "Cyathophyllum" ananas Goldfuss (1826); genus also includes "Pseudoacervularia" dybowskii Różkowska (1953), "Pseudoacervularia" intercellulosa (Phillips, 1841) sensu Pickett (1967), "Schlüteria" lyskovensis Ermakova (1957), "Acervularia" macrommata Roemer (1855), "Phillipsastrea" plantana Różkowska (1979), "Cyathophyllum" profundum Michelin (1845), "Phillipsastrea" rozkowskae Scrutton (1968), "Pseudoacervularia" cf. smithi (Różkowska, 1953) sensu Pickett (1967), "Phillipsastrea" ananas veserensis Coen-Aubert (1974) and "Phillipsastrea" zerda Galle, 1992 in Hladil et al. (1992).

Preisingerella[21]

Gen. et sp. nov

Valid

Kossovaya, Novak & Weyer

Early Permian

 Slovenia

A coral similar to members of the genus Caninia. The type species is P. stegovnikensis.

Ptychophyllum variatum[4]

Sp. nov

Valid

Munson & Jell

Silurian (late Gorstian to Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Ptychophyllidae.

Pycnostylus polyphyllodus[4]

Sp. nov

Valid

Munson & Jell

Silurian (late Wenlock)

Jack Formation

 Australia

A rugose coral belonging to the family Pycnostylidae.

Sokolovia[22]

Gen. et sp. nov

Junior homonym

Tsyganko

Late Devonian (Famennian)

 Russia
( Komi Republic)

A tabulate coral belonging to the family Pachyporidae. The type species is Sokolovia pershinae. The generic name is preoccupied by Sokolovia Ilovaisky (1934) and Sokolovia Shishkinskaya (1964).

Spinaxon[23]

Gen. et sp. nov

Valid

Weyer

Devonian (late Frasnian)

 France
 Germany

A member of Rugosa belonging to the family Petraiidae. The type species is Spinaxon potyi.

Syringopora hilarowiczi[19]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Syringoporidae; a species of Syringopora.

Syringopora kowalensis[19]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Syringoporidae; a species of Syringopora.

Talfania[24]

Gen. et sp. nov

Valid

Peel & McDermott

Ordovician (Katian)

Sholeshook Limestone Formation

 United Kingdom

A solitary coral. The type species is Talfania calicula.

Thamnoptychia mistiaeni[19]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Pachyporidae; a species of Thamnoptychia.

Utaratuia yunnanensis[10]

Sp. nov

Valid

Liao & Ma

Devonian (Eifelian)

 China

A rugose coral.

Vassiljukia[25]

Gen. et comb. nov

Valid

Denayer & Ogar

Carboniferous (Viséan)

 Turkey
 Ukraine

A colonial rugose coral; a new genus for "Lithostrotion" columnariformis Vassiljuk (1960).

Vesicospina[4]

Gen. et sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Kyphophyllidae. The type species is V. julli.

Vulykhia[26]

Nom. nov

Valid

Doweld

Late Ordovician

 Mongolia

An anthozoan belonging to the superfamily Heliolitoidea; a replacement name for Concavites Bondarenko & Minzhin (1981).

Arthropods

Bryozoans

Name Novelty Status Authors Age Unit Location Notes Images

Anisotrypa kjarkiensis[27]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Armenia

A bryozoan belonging to the group Trepostomata and the family Anisotrypidae.

Beisselina skyscanica[28]

Sp. nov

Valid

Koromyslova & Pakhnevich

Late Cretaceous (Campanian)

 Belarus

A bryozoan belonging to the group Ascophora and the family Tessaradomidae; a species of Beisselina.

Celleporaria pirabasensis[29]

Sp. nov

Valid

Muricy et al.

Oligocene-Miocene

Pirabas Formation

 Brazil

A bryozoan belonging to the group Cheilostomata and the family Lepraliellidae.

Celleporaria triangulavicularis[29]

Sp. nov

Valid

Muricy et al.

Oligocene-Miocene

Pirabas Formation

 Brazil

A bryozoan belonging to the group Cheilostomata and the family Lepraliellidae.

Dybowskiella hupehensiformis[30]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cystoporata belonging to the family Fistuliporidae.

Dyscritella lii[30]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Trepostomata belonging to the family Dystritellidae.

Ehrhardina[31]

Gen. et 2 sp. nov

Valid[32]

Martha & Taylor

Late Cretaceous (Cenomanian)

 France
 Germany
 United Kingdom

A bryozoan belonging to the group Flustrina and the family Onychocellidae. The type species is Ehrhardina voigti; genus also includes Ehrhardina pikeae.

Eridotrypella danzikensis[33]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Eridotrypellidae.

Etherella tibetensis[30]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cystoporata belonging to the family Etherellidae.

Fistuliphragma moniliformis[34]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistuliphragma sibirica[34]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistulipora sakagamii[30]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China
 Thailand

A member of Cystoporata belonging to the family Fistuliporidae.

Fistulipora salairiensis[34]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistuliramus fasciculus[34]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistulocladia cincinnata[34]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Metastenodiscus[35]

Gen. et comb. nov

Valid

Ernst, Schäfer & Grant-Mackie

Late Triassic

 New Caledonia
 New Zealand

A trepostome bryozoan. A new genus for "Stenodiscus" zealandicus Schäfer & Grant-Mackie (1994) and "Stenodiscus" kawhiae Schäfer & Grant-Mackie (1994).

Nikiforopora arpaensis[27]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Stenoporidae.

Pachydermopora grodnoensis[28]

Sp. nov

Valid

Koromyslova & Pakhnevich

Late Cretaceous (Campanian)

 Belarus

A bryozoan belonging to the group Ascophora and the family Tessaradomidae; a species of Pachydermopora.

Petalotrypa myunkhbalaensis[33]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata.

Planicellaria walsariensis[36]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Protoretepora irregularis[30]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Fenestrata belonging to the family Polyporidae.

Pyriporella charopadiensis[36]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Pyriporella vadsariensis[36]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Rectifenestella famenniensis[33]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the family Fenestellidae.

Rectifenestella kadrluiensis[33]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Armenia

A bryozoan belonging to the family Fenestellidae.

Stenophragmidium buckhornensis[37]

Sp. nov

Valid

Ernst et al.

Carboniferous (Pennsylvanian)

Boggy Formation

 United States
( Oklahoma)

Streblotrypa (Streblotrypa) heltzelae[37]

Sp. nov

Valid

Ernst et al.

Carboniferous (Pennsylvanian)

Boggy Formation

 United States
( Oklahoma)

Streblotrypa (Streblotrypa) parviformis[30]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cryptostomata belonging to the family Hyphasmoporidae.

Tabuliporella nakhichevanica[27]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Crustoporidae.

Tibetiporella[30]

Gen. et sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Fenestrata belonging to the family Polyporidae. The type species is T. ornata.

Timanotrypa australis[30]

Sp. nov

Valid

Ernst

Permian

Noonkanbah Formation
Rat Buri Limestone
Zongba Formation

 Australia
 China
 Thailand

A member of Cryptostomata belonging to the family Timanodictyidae.

Zigzagopora[38]

Gen. et sp. nov

Valid

Wilson & Taylor

Ordovician (Sandbian)

Bromide Formation

 United States
( Oklahoma)

A cyclostome bryozoan. Genus includes new species Z. wigleyensis.

Brachiopods

Name Novelty Status Authors Age Unit Location Notes Images

Acritosia ogamensis[39]

Sp. nov

Valid

Tazawa et al.

Early Permian (Kungurian)

Nabeyama Formation

 Japan

Acrosaccus scutatus[40]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Discinidae.

Anathyris (Anathyris) calestiennensis[41]

Sp. nov

Valid

Mottequin et al.

Devonian (Frasnian)

Nismes Formation

 Belgium

Anisopleurella antiqua[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Sowerbyellidae.

Apatobolus anoskelidion[40]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Obolidae.

Atansoria australis[40]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Obolidae.

Atryparia (Costatrypa) agricolae[43]

Sp. nov

Valid

Halamski & Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Atrypidae.

Aulacothyris waikatoensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Bellimurina fluctuosa[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Strophomenidae.

Biernatia pseudoplana[40]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Biernatiidae.

Biernatia wrighti[40]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Biernatiidae.

Biernatium minus[43]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Mystrophoridae.

Buxtonia inexpletucosta[45]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Canalilatus musculosus[46]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae.

Cerasinella[47]

Nom. nov

Valid

Copper

Silurian (Llandovery)

Merrimack Formation

 Canada
( Quebec)

An atrypoid brachiopod; a replacement name for Cerasina Copper (1995) (preoccupied).

Chapinella belkovskensis[48]

Sp. nov

Valid

Baranov, Sokiran & Blodgett

Devonian (Famennian)

Nerpalakhsk Formation

 Russia
( Sakha Republic)

A member of Rhynchonellida belonging to the family Pugnacidae.

Chilcatreta[49]

Gen. et sp. nov

Valid

Lavié & Benedetto

Ordovician (Darriwilian)

San Juan Formation

 Argentina

A siphonotretid brachiopod. Genus includes new species C. tubulata.

Crispithyris[44]

Gen. et sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida. Genus includes new species C. nauarchus.

Davidsonia enmerkaris[43]

Sp. nov

Valid

Halamski in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Davidsoniidae.

Dictyoclostus transversum[45]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Disculina mancenidoi[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Dolerorthis nadruvensis[50]

Sp. nov

Valid

Paškevičius & Hints

Ordovician (Katian)

 Lithuania

A member of Orthida belonging to the family Hesperorthidae.

Dulankarella hyrcanica[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Dyoros (Lissosia) maya[51]

Sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (Leonardian)

Paso Hondo Formation

 Mexico

Echinocoelia parva[43]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Ambocoeliidae.

Eochonetes maearum[52]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

Eochonetes minerva[52]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

Eochonetes voldemortus[52]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

Eoconulus puteus[40]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Eoconulidae.

Experilingula larga[46]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A member of Obolidae.

Flexaria magna[45]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Globiella kamiyassensis[53]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Glossella cuyanica[49]

Sp. nov

Valid

Lavié & Benedetto

Ordovician (Darriwilian)

San Juan Formation

 Argentina

Gondwanorthis[54]

Gen. et comb. nov

Valid[55]

Benedetto & Muñoz

Early Ordovician

 Argentina
 Iran

A plectorthoid brachiopod. A new genus for "Nanorthis" calderensis Benedetto (2007); genus also includes "Nanorthis" bastamensis Ghobadi Pour, Kebriaee-Zadeh & Popov (2011).

Gowanella[56]

Gen. et sp. nov

Valid

Hiller

Late Cretaceous (probably Maastrichtian)

Broken River Formation

 New Zealand

A member of Terebratulida related to Ostreathyris. The type species is G. capralis.

Grandispirifer qaidamensis[57]

Sp. nov

Valid

Lee, Shi & Chen in Shi et al.

Carboniferous (Serpukhovian)

Huaitoulata Formation

 China

A member of Spiriferoidea belonging to the family Spiriferidae.

Gundaria[58]

Gen. et sp. nov

Valid

Angiolini et al.

Permian

 Tajikistan

The type species is G. insolita.

Harperoides[59]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

 United States
( Alaska)

A member of Strophomenida belonging to the subfamily Mesodouvillininae. The type species is Harperoides alaskensis.

Hemileurus politus[58]

Sp. nov

Valid

Angiolini et al.

Permian

 Tajikistan

Holcothyris campbelli[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Hustedia shumardi[51]

Sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (Leonardian)

Paso Hondo Formation

 Mexico

Inflatia coodzavuii[45]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (late Mississippian, middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Iridistrophia (Flabellistrophia)[60]

Subgen. et comb. sp. nov

Valid

Jansen

Devonian (late Emsian to Eifelian)

 Germany
 Venezuela?

A member of Chilidiopsidae; a subgenus of Iridistrophia. The type species is "Orthis" hipponyx Schnur (1851); the subgenus also includes new species Iridistrophia (Flabellistrophia) musculosa and possibly also "Orthis" undifera Schnur (1853) and Iridistrophia dendritica Benedetto (1984).

Ishimia inflata[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Isogramma nakamurai[53]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Jakutoproductus lenensis[61]

Sp. nov

Valid

Makoshin

Early Permian

 Russia

Jakutoproductus talchanensis[61]

Sp. nov

Valid

Makoshin

Early Permian

 Russia

Kjaerina (Kjaerina) gondwanensis[62]

Sp. nov

Valid

Colmenar

Ordovician (late SandbianKatian)

Gabian Formation
Glauzy Formation
Louredo Formation
Portixeddu Formation

 France
 Italy
 Portugal

A rafinesquinid strophomenid brachiopod, a species of Kjaerina.

Kjaerina (Villasina)[62]

Subgen. et 3 sp. et comb. nov

Valid

Colmenar

Ordovician (Katian)

Cavá Formation
Gabian Formation
Portilla de Luna Limestones
Portixeddu Formation
Porto de Santa Anna Formation
Punta Serpeddi Formation
Rosan Formation

 France
 Italy
 Portugal
 Spain

A rafinesquinid strophomenid brachiopod, a subgenus of Kjaerina. The type species of the subgenus is Kjaerina (Villasina) pedronaensis; the subgenus also contains "Hedstroemina" almadenensis Villas (1995), as well as new species Kjaerina (Villasina) meloui and Kjaerina (Villasina) pyrenaica.

Koneviella? fuscina[46]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae.

Kutchithyris challinori[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Kutchithyris waitomoensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Lacunites jaroslavi[63]

Sp. nov

Valid

Mergl & Kraft

Early Ordovician

Klabava Formation

 Czech Republic

A paterinate brachiopod.

Lampazarorthis[54]

Gen. et comb. et sp. nov

Valid[55]

Benedetto & Muñoz

Early Ordovician

 Argentina

A plectorthoid brachiopod. A new genus for "Eoorthis" bifurcata Harrington (1937); genus also includes "Nanorthis" brachymyaria Benedetto in Benedetto & Carrasco (2002), as well as new species Lampazarorthis alata.

Lepidomena multiplicata[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Leptathyris gornensis[43]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Athyrididae.

Liaous[64]

Gen. et sp. nov

Valid

He & Chen in He et al.

Middle Triassic (early Anisian)

Xinyuan Formation

 China

A relative of Mentzelia and Paramentzelia. The type species is Liaous shaiwensis.

Loboidothyris awakinoensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Loboidothyris grantmackiei[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Loboidothyris marokopaensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Lyonia rochacamposi[65]

Sp. nov

Valid

Taboada et al.

Early Permian (latest Asselian-earliest Sakmarian)

Taciba Formation

 Brazil

A member of Productida belonging to the family Auriculispinidae, a species of Lyonia.

Mendozotreta[66]

Gen. et comb. nov

Valid

Holmer et al.

Ordovician

Antelope Valley Limestone
Lindero Formation

 Argentina
 United States
( Nevada)

A member of Acrotretida belonging to the family Acrotretidae. The type species is "Conotreta" devota Krause & Rowell (1975).

Mesoleptostrophia belli[67]

Sp. nov

Valid

Earp

Early Devonian

Montys Hut Formation

 Australia

Minutella bulgarica[68]

Sp. nov

Valid

Bitner & Motchurova-Dekova

Miocene (Badenian)

 Bulgaria

Monelasmina montisjosephi[43]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Draboviidae.

Nasakia[69]

Gen. et sp. nov

Valid

Streng et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Nasakia thulensis.

Numericoma rowelli[66]

Sp. nov

Valid

Holmer et al.

Ordovician (Darriwilian)

Antelope Valley Limestone
Ponon Trehue Formation

 Argentina
 United States
( Nevada)

A member of Lingulata belonging to the family Ephippelasmatidae.

Nushbiella kleithria[40]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Siphonotretidae.

Orthis dehmollaensis[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Orthidae.

Paraspirifer (Laurentispirifer)[60]

Subgen. et comb. nov

Valid

Jansen

Middle Devonian

 United States
 Venezuela

A subgenus of Paraspirifer. The type species is Paraspirifer conradi Godefroid & Fagerstrom (1983); the subgenus also includes "Delthyris" acuminata Conrad (1839), "Terebratula" acuminatissima de Castelnau (1843), "Spirifer" bownockeri Stewart (1927), Paraspirifer halli Godefroid & Fagerstrom (1983) and Paraspirifer clarkei Godefroid & Fagerstrom (1983).

Paraspirifer (Mosellospirifer)[60]

Subgen. et comb. nov

Valid

Jansen

Devonian (late Emsian to early Eifelian

 Germany
 Canada?
 China?

A subgenus of Paraspirifer. The type species is Paraspirifer sandbergeri Solle (1971); the subgenus also includes Spirifer auriculatus Sandberger & Sandberger (1856), Paraspirifer sandbergeri longimargo Solle (1971) (elevated to species rank), Paraspirifer eos Solle (1971) and Paraspirifer sandbergeri nepos Solle (1971). The subgenus might also include Paraspirifer gigantea Su (1976) and Paraspirifer desbiensi Bizzarro & Lespérance (1999).

Parazhanatella[46]

Gen. et sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae. The type species is P. paibia.

Pedderia[59]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

 United States
( Alaska)

A member of Rhynchonellida belonging to the family Pygmaellidae. The type species is Pedderia fragosa.

Phragmorthis shahrudensis[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Phragmorthidae.

Prospira pseudostruniana[70]

Sp. nov

Valid

Mottequin & Brice

Devonian (late Famennian)

Etrœungt Formation

 France

A member of Spiriferida belonging to the family Spiriferidae.

Psygmakantha[40]

Gen. et sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Ephippelasmatidae. The type species is P. malachiensis'.

Qaidamospirifer[57]

Gen. et sp. nov

Valid

Chen, Lee & Shi in Shi et al.

Carboniferous (Serpukhovian)

Huaitoulata Formation

 China

A member of Spiriferoidea belonging to the family Choristitidae. The type species is Q. elongatus.

Rafinesquina (Mesogeina)[62]

Subgen. et comb. et 2 sp. nov

Valid

Colmenar

Ordovician (Katian)

Bohdalec Formation
Fombuena Formation
Gabian Formation
Lower Ktaoua Formation
Porto de Santa Anna Formation
Upper Tiouririne Formation
Zahorany Formation

 Czech Republic
 France
 Germany
 Morocco
 Portugal
 Spain

A rafinesquinid strophomenid brachiopod, a subgenus of Rafinesquina. The type species of the subgenus is "Leptaena" pseudoloricata Barrande (1848); the subgenus also includes Rafinesquina pomoides Havlíček (1971), as well as new species Rafinesquina (Mesogeina) gabianensis and Rafinesquina (Mesogeina) loredensis.

Rhipidomella magna[53]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Rogorthis? oriens[42]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Orthidae.

Sampo suduvensis[50]

Sp. nov

Valid

Paškevičius & Hints

Late Ordovician

 Lithuania
 Russia
( Kaliningrad Oblast)

A member of Strophomenida belonging to the family Leptestiidae.

Sartenaerirhynchus[60]

Gen. et comb. nov

Valid

Jansen

Devonian (middle Siegenian to late Emsian)

 Belgium
 Germany
 Luxembourg

A member of Rhynchonellida belonging to the superfamily Uncinuloidea. The type species is "Terebratula" antiqua Schnur (1853); genus also includes "Uncinulus" frontecostatus Drevermann (1902)

Scaphelasma quadratum[40]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Scaphelasmatidae.

Siberioproductus[48]

Gen. et sp. nov

Valid

Baranov, Sokiran & Blodgett

Devonian (Famennian)

Chekursk Formation

 Russia
( Sakha Republic)

A member of Rhynchonelliformea belonging to the group Productida and the family Productellidae. The type species is S. boreus.

Skenidioides cretus[43]

Sp. nov

Valid

Halamski in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Skenidiidae.

Spinobolus[71]

Gen. et sp. nov

Valid

Zhang et al.

Cambrian

Shuijingtuo Formation

 China

A linguloid brachiopod. The type species is Spinobolus popovi.

Tapongaspirifer[67]

Gen. et sp. nov

Valid

Earp

Early Devonian

Montys Hut Formation

 Australia

A member of Spiriferida belonging to the family Filispiriferidae. The type species is Tapongaspirifer melodiae.

Taungurungetes[67]

Gen. et comb. nov

Valid

Earp

Early Devonian

Montys Hut Formation
Norton Gully Sandstone

 Australia

A possible member of Devonochonetinae. The type species is "Chonetes" taggertyensis Gill (1945) from the Montys Hut Formation; genus also contains second, unnamed species from the Norton Gully Sandstone.

Terebratulina leeae[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Terebratulina putiensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Thulatrypa[72]

Gen. et sp. et comb. nov

Valid

Huang et al.

Silurian

 China
 Norway

An atrypoid brachiopod. The type species is Thulatrypa gregaria; genus also contains "Meifodia" orientalis Rong, Xu & Yang (1974).

Tomteluva[69]

Gen. et sp. nov

Valid

Streng et al.

Cambrian

Stephen Formation

 Canada
( British Columbia)

A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Tomteluva perturbata.

Trentingula[73]

Gen. et 3 sp. et comb. nov

Valid

Posenato

Late Permian and Early Triassic

 Hungary
 Italy
 United States
( Wyoming)

A member of Linguloidea belonging to the family Lingulidae. The type species is T. lorigae; genus also includes new species T. mazzinensis and T. prinothi, as well as Trentingula borealis (Bittner, 1899).

Triangulospirifer[57]

Nom. nov

Valid

Lee in Shi et al.

Carboniferous

 United Kingdom

A member of Spiriferida. A replacement name for Triangularia Poletaev (2001).

Weberproductus[45]

Gen. et sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea. Genus includes new species W. donajiae.

Zeilleria opuatiaensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Zeilleria waiohipaensis[44]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Molluscs

Echinoderms

Name Novelty Status Authors Age Unit Location Notes Images

Alternocidaris[74]

Gen. nov

In press

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

 Egypt

A sea urchin.

Applinocrinus russelli[75]

Sp. nov

Valid[76]

Gale

Late Cretaceous (Maastrichtian)

Prairie Bluff Formation

 United States
( Mississippi)

A crinoid belonging to the group Articulata and the family Saccocomidae.

Apsidocrinus doreckae[77]

Sp. nov

Valid

Konieczyński, Pisera & Fózy

Early Cretaceous

 Hungary

A cyrtocrinid crinoid, a species of Apsidocrinus.

Arabicodiadema[78]

Gen. et comb. et sp. nov

Valid

Abdelhamid, El Qot & Abdelghany

Cretaceous (Albian to Cenomanian)

 Oman
 United Arab Emirates

A heterodiadematid sea urchin. The type species is "Trochodiadema" dhofarense Roman (1991); genus also includes new species Arabicodiadema alii.

Atalopegaster[79]

Gen. et sp. nov

Valid

Blake & Guensburg

Late Jurassic (Oxfordian)

Swift Formation

 United States
( Montana)

A starfish belonging to the family Stichasteridae. The type species is A. gundersoni.

Bystrowicrinus (col.) depressus[80]

Sp. nov

Valid

Donovan & Keighley

Late Silurian

West Point Formation

 Canada
( Quebec)

A crinoid.

Costatocrinus[75]

Gen. et 2 sp. nov

Valid[76]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Saccocomidae. The type species is C. brydonei; genus also includes C. mortimorei.

Cultellacrinus[75]

Gen. et sp. nov

Valid[76]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is C. gladius.

Darwinaster[81]

Gen. et sp. nov

Valid

Hunter, Rushton & Stone

Early Devonian

Fox Bay Formation

 Falkland Islands

A brittle star related to members of the genus Protaster. Genus includes new species D. coleenbiggsae.

Glyptocrinus nodosus[82]

Sp. nov

Valid

Kallmeyer & Ausich

Ordovician (Katian)

Kope Formation

 United States
( Kentucky
 Ohio)

A camerate crinoid, a species of Glyptocrinus.

Goniopygus macrotuberculatus[74]

Sp. nov

In press

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

 Egypt

A sea urchin.

Goniopygus subaequalis[74]

Sp. nov

In press

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

 Egypt

A sea urchin.

Hessicrinus[75]

Gen. et 2 sp. nov

Valid[76]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is H. filigree; genus also includes H. scalaensis.

Hiiumaacrinus[83]

Gen. et sp. nov

Valid

Ausich & Wilson

Silurian (Rhuddanian)

Hilliste Formation

 Estonia

A crinoid. Genus includes new species H. vinni.

Jakeocrinus[75]

Gen. et sp. nov

Valid[76]

Gale

Late Cretaceous (early Campanian)

Taylor Formation

 United States
( Texas)

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is J. ellisensis.

Lakotacrinus[84]

Gen. et sp. nov

Valid

Hunter et al.

Late Cretaceous (Campanian)

Pierre Shale

 United States
( South Dakota)

A stalked crinoid. The type species is L. brezinai.

Lehmannaster[85]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician

Traveusot Formation

 France

A stenuroid asterozoan. Genus includes new species L. spinosus.

Lopidiaster[85]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician

Garden City Formation

 United States
( Utah)

A stenuroid asterozoan. Genus includes new species L. jamisoni.

Loriolidiadema[78]

Gen. et comb. nov

Valid

Abdelhamid, El Qot & Abdelghany

Cretaceous (Aptian-Cenomanian)

 Egypt
 Portugal
Border region between  Syria and  Lebanon

A heterodiadematid sea urchin. The type species is "Pseudodiadema" libanoticum de Loriol (1887); genus also includes Loriolidiadema sculptile (de Loriol, 1887).

Lucernacrinus[75]

Gen. et sp. nov

Valid[76]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is L. woodi.

Maennilocystis[86]

Gen. et sp. nov

Valid

Paul & Rozhnov

Late Ordovician

 Estonia

A cystoid belonging to the family Callocystitidae. The type species is M. heckeri.

Mariania comaschicariae[87]

Sp. nov

Valid

Stara, Borghi & Kroh

Miocene (Aquitanian to early Burdigalian)

Nurallao Formation

 Italy

A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania.

Mariania stefaninii[87]

Sp. nov

Valid

Stara, Borghi & Kroh

Miocene (late Burdigalian to early Langhian)

Pantano Formation

 Italy

A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania.

Meturaster[85]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician

Lehman Formation

 United States
( Nevada)

A starfish. Genus includes new species M. belli.

Monophoraster telfordi[88]

Sp. nov

Valid

Mooi, Martínez & del Río

Early Miocene

Chenque Formation

 Argentina

A sand dollar.

Osteocrinus sinensis[89]

Sp. nov

Valid

Hess, Etter & Hagdorn

Late Triassic (early Carnian)

Xiaowa Formation

 China

A crinoid belonging to the group Roveacrinida, a species of Osteocrinus.

Pentamerocrinus kosovi[90]

Sp. nov

Valid

Rozhnov

Ordovician

 Russia
( Leningrad Oblast)

A crinoid belonging to the group Inadunata, a species of Pentamerocrinus.

Petraster crozonensis[85]

Sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician

Kermeur Formation

 France

A starfish.

Phyllocrinus bersekensis[77]

Sp. nov

Valid

Konieczyński, Pisera & Fózy

Early Cretaceous

 Hungary

A cyrtocrinid crinoid, a species of Phyllocrinus.

Placentinechinus[91]

Gen. et sp. nov

Valid[92]

Borghi & Garilli

Pleistocene (GelasianCalabrian)

 Italy

A sea urchin belonging to the family Temnopleuridae. The type species is P. davolii.

Protaeropsis[93]

Nom. nov

Valid

Doweld

Paleocene (Thanetian)

 Spain

A heart urchin; a replacement name for Sphenaster Jeffery in Smith et al. (1999).

Protaxocrinus estoniensis[83]

Sp. nov

Valid

Ausich & Wilson

Silurian (Rhuddanian)

Hilliste Formation

 Estonia

A crinoid.

Sagittacrinus[75]

Gen. et sp. nov

Valid[76]

Gale

Late Cretaceous (early Campanian)

Taylor Formation

 United States
( Texas)

A crinoid belonging to the group Articulata and the family Saccocomidae. The type species is S. torpedo.

Schuchertia sprinklei[85]

Sp. nov

Valid

Blake, Guensburg & Lefebvre

A starfish.

Stellacrinus[75]

Gen. et 2 sp. nov

Valid[76]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is S. hugesae; genus also includes S. pannosus.

Thinocrinus akanthos[94]

Sp. nov

Valid

Rhenberg, Ausich & Meyer

Carboniferous (Mississippian)

Fort Payne Formation

 United States

An actinocrinitid crinoid.

Conodonts

Name Novelty Status Authors Age Unit Location Notes Images

Amydrotaxis corniculans truncus[95]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Eognathodus kuangi[96]

Sp. nov

Valid

Earp

Early Devonian

 China

Eognathodus sulcatus lanei[95]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Eognathodus sulcatus sicatus[95]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Gladigondolella okayi[97]

Sp. nov

Valid

Kiliç

Triassic

Kayabaşı Limestone

 Turkey

Heliagnathus[95]

Gen. et sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida of uncertain phylogenetic placement. The type species is H. parvilabiatus.

Icriodus ampliatus[95]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Prioniodontida belonging to the family Icriodontidae.

Icriodus michiganus[98]

Sp. nov

Valid

Narkiewicz & Bultynck

Devonian (Eifelian)

Michigan Basin

 United States

Icriodus praealternatus ferus[99]

Subsp. nov

Valid

Wang et al.

Late Devonian (Frasnian/Famennian boundary)

Hongguleleng Formation

 China

A subspecies of Icriodus praealternatus.

Icriodus plurinodosus[99]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A species of Icriodus.

Icriodus stenoancylus junggarensis[99]

Subsp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A subspecies of Icriodus stenoancylus.

Idiognathodus lateralis[100]

Sp. nov

Valid

Hogancamp, Barrick & Strauss

Carboniferous (early Gzhelian)

 China
 Russia
 Ukraine
 United States

A member of Ozarkodinida belonging to the family Idiognathodontidae, a species of Idiognathodus.

Neogondolella cuneiforme[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)

Neogondolella curva[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella dilacerata[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)

Neogondolella hastata[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella indicta[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella panlaurentia[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella tenera[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella vellicata[101]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neopolygnathus huijunae[99]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A species of Neopolygnathus.

Neospathodus planus[102]

Sp. nov

Valid

Chen et al.

Early Triassic

 Slovenia

Oulodus astriatus[95]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Prioniodinida belonging to the family Prioniodinidae.

Ozarkodina multistriola[95]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina scoliciformis[95]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi cordata[95]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi lanceola[95]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi lenticula[95]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Panderodus rhytiodus[95]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of the family Panderodontidae.

Pelekysgnathus inequalis[95]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Prioniodontida belonging to the family Icriodontidae.

Pelekysgnathus jeppssoni[103]

Sp. nov

Valid

Nazarova

Devonian (Eifelian)

 Russia
( Bryansk Oblast)

Platyvillosus corniger[102]

Sp. nov

Valid

Chen et al.

Early Triassic

 Slovenia

Polygnathus abaimovae[104]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus beckeri[104]

Nom. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

A replacement name for Polygnathus inflexus Baranov (1992).

Polygnathus datnensis[104]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus dogdensis[104]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus novozemelicus[104]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

“Polygnathus” pseudocommunis[99]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

Possibly a species of Polygnathus.

Polygnathus slastenovi[104]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus tarabukini[104]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Staeschegnathus[105]

Gen. et sp. nov

Valid

Koike

Triassic

Taho Formation

 Japan

An ellisonid conodont. The type species is S. perrii.

Fishes

Amphibians

Research

New taxa

Temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Cyclotosaurus buechneri[114]

Sp. nov

Valid

Witzmann, Sachs & Nyhuis

Late Triassic (middle Carnian)

Stuttgart Formation

 Germany

A mastodonsauroid temnospondyl, a species of Cyclotosaurus.

Konzhukovia sangabrielensis[115]

Sp. nov

Valid[116]

Pacheco et al.

Permian (early Guadalupian)

Rio do Rasto Formation

 Brazil

Samarabatrachus[117]

Gen. et sp. nov

Valid

Novikov

Early Triassic

 Russia

The type species is S. bjerringi.

Selenocara rossica[117]

Sp. nov

Valid

Novikov

Early Triassic

 Russia

Stanocephalosaurus amenasensis[118]

Sp. nov

Valid

Dahoumane et al.

Early-Middle Triassic

Zarzaïtine Series, Illizi Basin

 Algeria

A mastodonsauroid temnospondyl, a species of Stanocephalosaurus.

Syrtosuchus[117]

Gen. et comb. et sp. nov

Valid

Novikov

Early Triassic

 Russia

Genus includes "Wetlugasaurus" samarensis Sennikov (1981), as well as new species S. morkovini.

Tomeia[119]

Gen. et sp. nov

Valid

Eltink, Da-Rosa & Dias-da-Silva

Early Triassic

Sanga do Cabral Supersequence

 Brazil

A mastodonsauroid temnospondyl. The type species is T. witecki.

Yuanansuchus maopingchangensis[120]

Sp. nov

Valid

Liu

Middle Triassic (Anisian)

Badong Formation

 China

A mastodonsauroid temnospondyl, a species of Yuanansuchus.

Lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Hyogobatrachus[121]

Gen. et sp. nov

Valid

Ikeda, Ota & Matsui

Early Cretaceous

Sasayama Group

 Japan

A frog. The type species is Hyogobatrachus wadai.

Kiyatriton krasnolutskii[122]

Sp. nov

Valid

Skutschas

Middle Jurassic (Bathonian)

Itat Formation

 Russia
( Krasnoyarsk Krai)

A salamander, a species of Kiyatriton.

Kuruleufenia[123]

Gen. et sp. nov

Valid

Gómez

Late Cretaceous (late Campanian–early Maastrichtian)

Allen Formation

 Argentina

A member of Pipidae. The type species is Kuruleufenia xenopoides.

Litoria lundeliusi[124]

Sp. nov

Valid

Tyler & Prideaux

Pleistocene

 Australia

A species of Litoria.

Nuominerpeton[125]

Gen. et sp. nov

Valid

Jia & Gao

Early Cretaceous (BarremianAptian)

Guanghua Formation

 China

A cryptobranchoid salamander of uncertain phylogenetic placement. The type species is N. aquilonaris.

Palaeobatrachus eurydices[126]

Sp. nov

Valid

Villa et al.

Early Pleistocene (Gelasian)

 Netherlands

Phosphotriton[127]

Gen. et sp. nov

Valid

Tissier, Rage, Boistel, Fernandez, Pollet, Garcia and Laurin

Eocene

 France

A salamander. The type species is Phosphotriton sigei.

Prospea[128]

Gen. et sp. nov

Valid

Chen et al.

Paleocene

Naranbulak Formation

 Mongolia

A member of Scaphiopodidae. The type species is Prospea holoserisca.

Qinglongtriton[129]

Gen. et sp. nov

Valid

Jia & Gao

Late Jurassic (Oxfordian)

Tiaojishan Formation

 China

A basal member of Salamandroidea. The type species is Qinglongtriton gangouensis.

Tambabatrachus[121]

Gen. et sp. nov

Valid

Ikeda, Ota & Matsui

Early Cretaceous

Sasayama Group

 Japan

A frog. The type species is Tambabatrachus kawazu.

Lepidosaurs

Lizards

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Bagaluus[138]

Gen. et sp. nov

Valid

Alifanov

Early Cretaceous

 Mongolia

A member of Scincomorpha belonging to the family Hodzhakuliidae. The type species is B. primigenius.

Carnoscincus[138]

Gen. et sp. nov

Valid

Alifanov

Early Cretaceous

 Mongolia

A member of Scincomorpha belonging to the family Hodzhakuliidae. The type species is C. eublepharus.

Janosikia[139]

Gen. et comb. nov

Valid

Čerňanský, Klembara & Smith

Early Miocene

 Germany

A member of Lacertidae; a new genus for "Ophisaurus" ulmensis Gerhardt (1903).

Jeddaherdan[140]

Gen. et sp. nov

Apesteguía et al.

Late Cretaceous (Cenomanian)

Kem Kem Beds

 Morocco

An iguanian belonging to the group Acrodonta, possibly a relative of the uromasticine agamids. The type species is J. aleadonta.

Ophisauromimus[141]

Gen. et comb. nov

Valid

Čerňanský, Klembara & Műller

Oligocene

 France
 Germany

A member of Anguidae. A new genus for "Dopasia" coderetensis Augé (2005); genus also includes "Dopasia" frayssensis Augé (2005).

Platynotoides[138]

Gen. et sp. nov

Junior homonym

Alifanov

Early Cretaceous

 Mongolia

A member of Scincomorpha belonging to the family Hodzhakuliidae. The type species is P. altidentatus. The generic name is preoccupied by Platynotoides Kaszab (1975).

Pluridens calabaria[142]

Sp. nov

Valid

Longrich

Late Cretaceous (late Campanian)

Nkporo Shale

 Nigeria

A mosasaur, a species of Pluridens.

Solastella[143]

Gen. et sp. nov

Valid

Stocker & Kirk

Eocene

Devil's Graveyard Formation

 United States
( Texas)

A rhineurid amphisbaenian. The type species is Solastella cookei.

Snakes

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Lunaophis[149]

Gen. et sp. nov

Valid

Albino, Carrillo-Briceño & Neenan

Late Cretaceous (Cenomanian)

La Luna Formation

 Venezuela

A snake of uncertain phylogenetic placement. The type species is L. aquaticus.

Platyspondylophis[150]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

Cambay Shale Formation

 India

A member of Madtsoiidae. The type species is P. tadkeshwarensis.

Rieppelophis[151]

Gen. et comb. nov

Valid

Scanferla, Smith & Schaal

Eocene

Messel pit

 Germany

A member of Boidae. A new genus for "Messelophis" ermannorum Schaal & Baszio (2004).

Ichthyosauromorphs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Cryopterygius kielanae[154]

Sp. nov

Valid

Tyborowski

Late Jurassic (Tithonian)

Kcynia Formation

 Poland

A member of Ophthalmosauridae.

Ichthyosaurus larkini[155]

Sp. nov

Valid[156]

Lomax & Massare

Early Jurassic (Hettangian)

 United Kingdom

Ichthyosaurus somersetensis[155]

Sp. nov

Valid[156]

Lomax & Massare

Early Jurassic (Hettangian)

 United Kingdom

Sclerocormus[157]

Gen. et sp. nov

Valid

Jiang et al.

Early Triassic (Olenekian)

Nanlinghu Formation

 China

A basal member of Ichthyosauriformes. The type species is S. parviceps.

Wahlisaurus[158]

Gen. et sp. nov

Valid[159]

Lomax

Early Jurassic (Hettangian)

 United Kingdom

A member of Leptonectidae. The type species is W. massarae.

Sauropterygians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Alexandronectes[163]

Gen. et sp. nov

Valid

Otero et al.

Late Cretaceous (early Maastrichtian)

Conway Formation

 New Zealand

An aristonectine elasmosaurid plesiosaur. The type species is Alexandronectes zealandiensis.

Dawazisaurus[164]

Gen. et sp. nov

Valid

Cheng et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A non-pistosauroid eosauropterygian of uncertain phylogenetic placement. The type species is Dawazisaurus brevis.

Kawanectes[165]

Gen. et comb. nov

Valid

O'Gorman

Late Cretaceous (late Campanian–early Maastrichtian)

Allen Formation

 Argentina

An elasmosaurid plesiosaur. The type species is "Trinacromerum" lafquenianum Gasparini & Goñi (1985).

Lariosaurus vosseveldensis[166]

Sp. nov

Valid

Klein et al.

Middle Triassic (Anisian)

 Netherlands

Polycotylus sopozkoi[167]

Sp. nov

Valid

Efimov, Meleshin & Nikiforov

Late Cretaceous

 Russia

Stenorhynchosaurus[168]

Gen. et sp. nov

Valid

Páramo et al.

Early Cretaceous (late Barremian)

Paja Formation

 Colombia

A pliosaurid plesiosaur. The type species is Stenorhynchosaurus munozi.

Turtles

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Algorachelus[177]

Gen. et sp. nov

Valid

Pérez-García[178]

Late Cretaceous (Cenomanian)

 Spain

A member of Bothremydidae. The type species is A. peregrinus.

Anhuichelys doumuensis[179]

Sp. nov

Valid

Tong et al.

Middle Paleocene

Doumu Formation

 China

A stem-tortoise, a species of Anhuichelys.

Fontainechelon[180]

Gen. et comb. nov

Valid

Pérez-García, Ortega & Jiménez Fuentes

Early Eocene

 France

A tortoise; a new genus for "Achilemys" cassouleti Claude & Tong (2004).

Inaechelys[181]

Gen. et sp. nov

Valid

Carvalho, Ghilardi & Barreto

Paleocene (Danian)

Maria Farinha Formation

 Brazil

A member of Bothremydidae. The type species is I. pernambucensis. Its status as a valid taxon was challenged by Romano (2016).[182]

Keuperotesta[183]

Gen. et sp. nov

Valid

Szczygielski & Sulej

Late Triassic

 Germany

A member of Proterochersidae. The type species is Keuperotesta limendorsa.

Kinosternon notolophus[184]

Sp. nov

Valid

Bourque

Miocene (Clarendonian)

Alachua Formation
Statenville Formation

 United States
( Florida)

A mud turtle.

Kinosternon pannekollops[184]

Sp. nov

Valid

Bourque

Miocene (Clarendonian)

Ogallala Formation

 United States
( Texas)

A mud turtle.

Kinosternon rincon[184]

Sp. nov

Valid

Bourque

Miocene (late Barstovian)

Cerro Conejo Formation

 United States
( New Mexico)

A mud turtle.

Kinosternon wakeeniense[184]

Sp. nov

Valid

Bourque

Miocene (Clarendonian)

Ash Hollow Formation
Ogallala Formation

 United States
( Kansas,
 Nebraska)

A mud turtle.

Neurankylus notos[185]

Sp. nov

Valid

Lichtig & Lucas

Late Cretaceous (Coniacian-Santonian)

Crevasse Canyon Formation

 United States
( New Mexico)

A member of Baenidae.

Neurankylus torrejonensis[186]

Sp. nov

Valid

Lyson et al.

Paleocene (Torrejonian)

Nacimiento Formation

 United States
( New Mexico)

A member of Baenidae.

Paiutemys[187]

Gen. et sp. nov

Valid

Joyce, Lyson & Kirkland

Late Cretaceous (late Cenomanian)

 United States
( Utah)

A member of Bothremydidae. The type species is P. tibert.

Palaeoamyda[188]

Gen. et comb. nov

Valid

Cadena

Eocene

 Germany

A relative of trionychids; a new genus for "Trionyx" messelianus Reinach (1900).

Pelorochelon[180]

Gen. et sp. et comb. nov

Valid

Pérez-García, Ortega & Jiménez Fuentes

Middle Eocene

 Germany
 Spain

A tortoise. The type species is P. soriana; genus also includes Pelorochelon eocaenica (Hummel, 1935).

Proterochersis porebensis[183]

Sp. nov

Valid

Szczygielski & Sulej

Late Triassic

 Poland

A member of Proterochersidae.

Sichuanchelys palatodentata[189]

Sp. nov

Valid

Joyce et al.

Late Jurassic (Oxfordian)

Shishugou Formation

 China

A basal member of Testudinata.

Tartaruscola[190]

Gen. et sp. nov

Valid

Pérez-García

Eocene (Ypresian)

 France

A member of Bothremydidae belonging to the group Foxemydina. The type species is T. teodorii.

Yelmochelys[191]

Gen. et sp. nov

Valid

Brinkman et al.

Late Cretaceous (late Campanian and early Maastrichtian)

Cañon del Tule Formation
Cerro del Pueblo Formation

 Mexico

A stem-kinosternid. The type species is Yelmochelys rosarioae.

Archosauriformes

Basal archosauriforms

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Litorosuchus[199]

Gen. et sp. nov

Valid

Li et al.

Middle Triassic

Falang Formation

 China

Probably a relative of Vancleavea. The type species is L. somnii.

Triopticus[200]

Gen. et sp. nov

Valid

Stocker et al.

Late Triassic (latest Carnian-early Norian)

Dockum Group

 United States
( Texas)

Probably a basal member of Archosauriformes. The type species is T. primus.

Pseudosuchians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Agaresuchus[222]

Gen. et sp. et comb. nov

Valid

Narváez et al.

Late Cretaceous (late CampanianMaastrichtian)

 Spain

A member of Allodaposuchidae. Genus includes new species Agaresuchus fontisensis, as well as Allodaposuchus” subjuniperus.

Bayomesasuchus[223]

Gen. et sp. nov

Valid

Barrios, Paulina-Carabajal & Bona

Late Cretaceous

Cerro Lisandro Formation

 Argentina

A peirosaurid crocodyliform. The type species is Bayomesasuchus hernandezi.

Elosuchus broinae[224]

Sp. nov

In press

Meunier & Larsson

Late Cretaceous (Cenomanian)

 Algeria

Fortignathus[225]

Gen. et comb. nov

Valid[226]

Young et al.

Cretaceous (late Albian-early Cenomanian)

Echkar Formation

 Niger

A dyrosaurid or a relative of dyrosaurids; a new genus for "Elosuchus" felixi de Lapparent de Broin (2002).

Gryposuchus pachakamue[227]

Sp. nov

Valid

Salas-Gismondi et al.

Miocene

Pebas Formation

 Peru

A member of Gryposuchinae, a species of Gryposuchus.

Kalthifrons[228]

Gen. et sp. nov

Valid

Yates & Pledge

Pliocene

Tirari Formation

 Australia

A member of Mekosuchinae. The type species is K. aurivellensis.

Kentisuchus astrei[229]

Sp. nov

Valid

Jouve

Eocene (late Lutetian)

 France

A member of Tomistominae, a species of Kentisuchus.

Lavocatchampsa[230]

Gen. et sp. nov

Valid

Martin & De Lapparent De Broin

Cretaceous (Albian-Cenomanian)

Kem Kem Beds

 Morocco

A notosuchian. The type species is L. sigogneaurusselae.

Llanosuchus[231]

Gen. et sp. nov

Valid

Fiorelli et al.

Late Cretaceous (Campanian?)

Los Llanos Formation

 Argentina

A notosuchian crocodyliform. The type species is Llanosuchus tamaensis.

Machimosaurus rex[232]

Sp. nov

Valid

Fanti et al.

Early Cretaceous

 Tunisia

A teleosaurid crocodylomorph, a species of Machimosaurus.

Patagosuchus[233]

Gen. et sp. nov

Valid

Lio et al.

Late Cretaceous (Turonian–Coniacian)

Portezuelo Formation

 Argentina

A peirosaurid crocodylomorph. The type species is Patagosuchus anielensis.

Protoalligator[234]

Gen. et comb. nov

Valid

Wang, Sullivan & Liu

Middle Paleocene

Wanghudun Formation

 China

A member of Alligatoroidea of uncertain phylogenetic placement; a new genus for "Eoalligator" huiningensis Young (1982).

Sabinosuchus[235]

Gen. et sp. nov

Valid

Shiller, Porras-Muzquiz & Lehman

Late Cretaceous (Maastrichtian)

Escondido Formation

 Mexico

A member of Dyrosauridae. The type species is S. coahuilensis.

Sabresuchus[236]

Gen. et comb. nov

Valid

Tennant, Mannion & Upchurch

Cretaceous (late BarremianMaastrichtian)

 Romania
 Spain

A member of Paralligatoridae. The type species is "Theriosuchus" ibericus Brinkmann (1989); genus also includes "Theriosuchus" sympiestodon Martin, Rabi & Csiki (2010).

Scutarx[237][238]

Gen. et sp. nov

Valid

Parker

Late Triassic (middle Norian)

Chinle Formation
Cooper Canyon Formation

 United States
( Arizona,  Texas)

An aetosaur. The type species is Scutarx deltatylus.

Ultrastenos[239]

Gen. et sp. nov

Valid

Stein, Hand & Archer

Late Oligocene

Riversleigh World Heritage Area

 Australia

A member of Mekosuchinae. The type species is U. willisi.

Vivaron[240]

Gen. et sp. nov

Valid

Lessner et al.

Late Triassic (Norian)

Chinle Formation

 United States
( New Mexico)

A rauisuchid. The type species is V. haydeni.

Basal dinosauromorphs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Dromomeron gigas[244]

Sp. nov

Valid

Martínez et al.

Late Triassic (Norian)

Quebrada del Barro Formation

 Argentina

A lagerpetid dinosauromorph, a species of Dromomeron.

Ixalerpeton[245]

Gen. et sp. nov

Valid

Cabreira et al.

Late Triassic (Carnian)

Santa Maria Formation

 Brazil

A lagerpetid dinosauromorph. The type species is I. polesinensis.

Non-avian dinosaurs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Agujaceratops mavericus[314]

Sp. nov

Valid[315]

Lehman, Wick & Barnes

Late Cretaceous

Aguja Formation

 United States
( Texas)

A chasmosaurine ceratopsian.

Alcovasaurus[316]

Gen. et comb. nov

Valid

Galton & Carpenter

Late Jurassic

Morrison Formation

 United States
( Wyoming)

A stegosaur; a new genus for "Stegosaurus" longispinus Gilmore (1914). This species was previously made the type species of the new genus Natronasaurus by Ulansky (2014); however, Galton & Carpenter (2016) claim it did not meet the requirements of the International Code of Zoological Nomenclature.[316]

Aoniraptor[317]

Gen. et sp. nov

Valid

Motta et al.

Late Cretaceous (middle Cenomanian-early Turonian)

Huincul Formation

 Argentina

A theropod dinosaur of uncertain phylogenetic placement, a possible relative of Deltadromeus. The type species is A. libertatem.

Apatoraptor[318]

Gen. et sp. nov

Valid

Funston & Currie

Late Cretaceous

Horseshoe Canyon Formation

 Canada
( Alberta)

A caenagnathid theropod. The type species is Apatoraptor pennatus.

Austroposeidon[319]

Gen. et sp. nov

Valid

Bandeira et al.

Late Cretaceous (Campanian-Maastrichtian)

Presidente Prudente Formation

 Brazil

A titanosaur sauropod. The type species is A. magnificus.

Beipiaognathus[320]

Gen. et sp. nov

Valid

Hu, Wang & Huang

Early Cretaceous

Yixian Formation

 China

A compsognathid theropod. The type species is B. jii.

Buriolestes[245]

Gen. et sp. nov

Valid

Cabreira et al.

Late Triassic (Carnian)

Santa Maria Formation

 Brazil

A basal member of Sauropodomorpha. The type species is B. schultzi.

Datonglong[321]

Gen. et sp. nov

Valid

Xu et al.

Late Cretaceous

Huiquanpu Formation

 China

A non-hadrosaurid hadrosauroid ornithopod. The type species is Datonglong tianzhenensis.

Dracoraptor[322]

Gen. et sp. nov

Valid

Martill et al.

Early Jurassic (Hettangian)

Blue Lias Formation

 United Kingdom

A basal member of Neotheropoda. The type species is Dracoraptor hanigani.

Eotrachodon[323][324]

Gen. et sp. nov

Valid

Prieto-Marquez, Erickson & Ebersole

Late Cretaceous (latest Santonian)

Mooreville Chalk

 United States
( Alabama)

A hadrosaurid ornithopod. The type species is Eotrachodon orientalis.

Foraminacephale[325]

Gen. et comb. nov

Valid

Schott & Evans

Late Cretaceous (Campanian)

 Canada
( Alberta)

A new genus for "Stegoceras" brevis Lambe (1918).

Fukuivenator[326]

Gen. et sp. nov

Valid

Azuma et al.

Early Cretaceous (Barremian or Aptian)

Kitadani Formation

 Japan

A member of Maniraptora of uncertain phylogenetic placement. The type species is Fukuivenator paradoxus.

Gastonia lorriemcwhinneyae[327]

Sp. nov

Valid

Kinneer, Carpenter & Shaw

Early Cretaceous

Cedar Mountain Formation

 United States
( Utah)

?Gryposaurus alsatei[328]

Sp. nov

Valid

Lehman, Wick & Wagner

Late Cretaceous (Maastrichtian)

Javelina Formation

 United States
( Texas)

A hadrosaurid, possibly a species of Gryposaurus.

Gualicho[329]

Gen. et sp. nov

Valid

Apesteguía et al.

Late Cretaceous (Cenomanian to Turonian)

Huincul Formation

 Argentina

A theropod dinosaur of uncertain phylogenetic placement, a possible relative of Deltadromeus. The taxon informally referred to as "Nototyrannus" before its formal description. The type species is G. shinyae.

Lohuecotitan[330]

Gen. et sp. nov

In press

Díaz et al.

Late Cretaceous (late Campanian-early Maastrichtian)

 Spain

A titanosaur sauropod. The type species is L. pandafilandi.

Machairoceratops[331]

Gen. et sp. nov

Valid

Lund et al.

Late Cretaceous (Campanian)

Wahweap Formation

 United States
( Utah)

A centrosaurine ceratopsian. The type species is Machairoceratops cronusi.

Meroktenos[332]

Gen. et comb. nov

Valid

Peyre de Fabrègues & Allain

Late Triassic

Lower Elliot Formation

 Lesotho

A non-sauropod sauropodomorph. The type species is "Melanorosaurus" thabanensis Gauffre (1993).

Morrosaurus[333]

Gen. et sp. nov

Valid

Rozadilla et al.

Late Cretaceous (Maastrichtian)

López de Bertodano Formation

 Antarctica

An iguanodontian ornithopod. The type species is Morrosaurus antarcticus.

Murusraptor[334]

Gen. et sp. nov

Valid

Coria & Currie

Late Cretaceous (Coniacian)

Sierra Barrosa Formation

 Argentina

A theropod belonging to the group Megaraptora. The type species is M. barrosaensis.

Notocolossus[335]

Gen. et sp. nov

Valid

González Riga et al.

Late Cretaceous (late Coniacian–early Santonian)

Plottier Formation

 Argentina

A titanosaur sauropod. The type species is Notocolossus gonzalezparejasi.

Rativates[336]

Gen. et sp. nov

Valid

McFeeters et al.

Late Cretaceous (late Campanian)

Dinosaur Park Formation

 Canada
( Alberta)

An ornithomimid theropod. The type species is R. evadens.

Sarmientosaurus[337]

Gen. et sp. nov

Valid

Martínez et al.

Late Cretaceous (Cenomanian-Turonian)

Bajo Barreal Formation

 Argentina

A titanosaur sauropod, a basal member of Lithostrotia. The type species is Sarmientosaurus musacchioi.

Savannasaurus[338]

Gen. et sp. nov

Poropat et al.

Late Cretaceous (Cenomanian-early Turonian)

Winton Formation

 Australia

A titanosaur sauropod. The type species is S. elliottorum.

Spiclypeus[339]

Gen. et sp. nov

Valid

Mallon et al.

Late Cretaceous (late Campanian)

Judith River Formation

 United States
( Montana)

A chasmosaurine ceratopsian. The type species is Spiclypeus shipporum.

Taurovenator[317]

Gen. et sp. nov

Valid

Motta et al.

Late Cretaceous (middle Cenomanian-early Turonian)

Huincul Formation

 Argentina

A carcharodontosaurid theropod. The type species is T. violantei.

Timurlengia[340]

Gen. et sp. nov

Valid

Brusatte et al.

Late Cretaceous (Turonian)

Bissekty Formation

 Uzbekistan

A non-tyrannosaurid tyrannosauroid. The type species is Timurlengia euotica.

Tongtianlong[341]

Gen. et sp. nov

et al.

Late Cretaceous (Maastrichtian)

Nanxiong Formation

 China

An oviraptorid theropod. The type species is T. limosus.

Tototlmimus[342]

Gen. et sp. nov

Valid

Serrano-Brañas et al.

Late Cretaceous

Packard Shale Formation

 Mexico

An ornithomimid theropod. The type species is Tototlmimus packardensis.

Viavenator[343]

Gen. et sp. nov

Valid

Filippi et al.

Late Cretaceous (Santonian)

Bajo de la Carpa Formation

 Argentina

A brachyrostran abelisaurid theropod. The type species is Viavenator exxoni.

Wiehenvenator [344]

Gen. et sp. nov.

Valid

Rauhut, Hübner & Lanser

Middle Jurassic (Callovian)

Ornatenton Formation

 Germany

A megalosaurid theropod. The type species is W. albati.

Birds

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Antarctoboenus [388]

Gen. et sp. nov.

Valid

Cenizo, Noriega & Reguero

Early Eocene

La Meseta Formation

 Antarctica

(Seymour Island)

A stem-falconid. The type species is A. carlinii.

Bellulornis [389][390]

Gen. et sp. nov.

Valid

Wang, Zhou & Zhou

Early Cretaceous

Jiufotang Formation

 China

A basal member of Ornithuromorpha. The type species is B. rectusunguis. The original generic name was Bellulia, which turned out to be preoccupied by Bellulia Fibiger (2008).

Calciavis [391]

Gen. et sp. nov.

Valid

Nesbitt & Clarke

Early Eocene

Green River Formation

 United States
( Wyoming)

A member of Lithornithidae. The type species is C. grandei.

Centropus bairdi [392]

Sp. nov.

Valid

Shute, Prideaux & Worthy

Pleistocene

 Australia

A member of the Cuculidae.

Centropus maximus [392]

Sp. nov.

Valid

Shute, Prideaux & Worthy

Pleistocene

 Australia

A member of the Cuculidae.

Changzuiornis [393]

Gen. et sp. nov.

Valid

Huang et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

An early member of Euornithes. The type species is C. ahgmi.

Chiappeavis [394]

Gen. et sp. nov.

Valid

O’Connor et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes, probably belonging to the family Pengornithidae. The type species is C. magnapremaxillo.

Chionoides [395]

Gen. et sp. nov.

Valid

De Pietri et al.

Late Oligocene

 Australia

A member of Chionoidea of uncertain phylogenetic placement, showing the mosaic of characters shared with both sheathbills and the Magellanic plover. The type species is C. australiensis.

Chongmingia [396]

Gen. et sp. nov.

Valid

Wang et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Avialae of uncertain phylogenetic placement. The type species is C. zhengi.

Cypseloramphus [397]

Gen. et sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

Possibly a basal member of Apodiformes. The type species is C. dimidius.

Daphoenositta trevorworthyi [398]

Sp. nov.

Valid

Nguyen

Miocene

Riversleigh World Heritage Area

 Australia

A sittella

Dingavis [399]

Gen. et sp. nov.

Valid [400]

O'Connor, Wang & Hu

Early Cretaceous

Yixian Formation

 China

A basal member of Ornithuromorpha. The type species is D. longimaxilla.

Dromornis murrayi [401]

Sp. nov.

Valid

Worthy et al.

Late Oligocene–Early Miocene

Riversleigh

 Australia

A member of Dromornithidae

Eostrix gulottai [402]

Sp. nov.

Valid

Mayr

Early Eocene

Nanjemoy Formation

 United States
( Virginia)

An early owl of the family Protostrigidae.

Eurobambusicola [378]

Gen. et sp. nov.

Valid

Zelenkov

Late Miocene

 Hungary

A member of the family Phasianidae. The type species is E. turolicus.

Galligeranoides [403]

Gen. et sp. nov.

Valid

Bourdon, Mourer-Chauviré, & Laurent

middle Ypresian

 France

A member of the family Geranoididae. The type species is G. boriensis.

Gallinago kakuki [404]

Sp. nov.

Valid

Steadman & Takano

Late Quaternary

 The Bahamas
 Cayman Islands
 Cuba

A member of Scolopacidae, a species of Gallinago.

Hesperornis lumgairi [405]

Sp. nov.

Valid

Aotsuka & Sato

Campanian

Pierre Shale

 Canada

A species of Hesperornis.

Klallamornis abyssa [406]

Gen. et sp. nov.

Valid

Mayr & Goedert

Latest Eocene or Early Oligocene

 United States
( Washington)

A member of Plotopteridae. This is the type species of the new genus.

?Klallamornis clarki [406]

Sp. nov.

Valid

Mayr & Goedert

Latest Eocene or Early Oligocene

 United States
( Washington)

A member of Plotopteridae. possibly a species of Klallamornis.

Lapillavis [397]

Gen. et sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

A bird of uncertain phylogenetic placement, showing similarities to Foshanornis songi. The type species is L. incubarens.

Linyiornis [407]

Gen. et sp. nov.

Valid

Wang et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes. The type species is L. amoena.

Mioneophron [408]

Gen. et sp. nov.

Valid

Li et al.

Late Miocene

Liushu Formation

 China

A member of Gypaetinae Vieillot (1816). The type species is M. longirostris.

Mioryaba [378]

Gen. et sp. nov.

Valid

Zelenkov

Late Miocene

 Hungary

A member of the family Phasianidae. The type species is M. magyarica.

Monoenantiornis [409]

Gen. et sp. nov.

Valid[410]

Hu & O’Connor

Early Cretaceous

Yixian Formation

 China

A member of Enantiornithes. The type species is M. sihedangia.

Neilus [395]

Gen. et sp. nov.

Valid

De Pietri et al.

Early Miocene

 New Zealand

A member of Chionoidea of uncertain phylogenetic placement, showing the mosaic of characters shared with both sheathbills and the Magellanic plover. The type species is N. sansomae.

Notoleptos [411]

Gen. et sp. nov

Valid

Acosta Hospitaleche & Gelfo

Late Eocene

 Antarctica

(Seymour Island)

A probable relative of albatrosses. The type species is N. giglii.

Olympidytes [406]

Gen. et sp. nov.

Valid

Mayr & Goedert

Latest Eocene or Early Oligocene

 United States
( Washington)

A member of Plotopteridae. The type species is O. thieli.

Phalcoboenus napieri [412]

Sp. nov.

Valid

Adams & Woods

Holocene

 Falkland Islands

A member of Phalcoboenus.

Primozygodactylus longibrachium [413]

Sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

A member of Zygodactylidae.

Primozygodactylus quintus [413]

Sp. nov.

Valid

Mayr

Early Eocene

Messel pit

 Germany

A member of Zygodactylidae.

Protomelanitta bakeri [414]

Sp. nov.

Valid

Stidham & Zelenkov

Miocene

Esmeralda Formation

 United States
( Nevada)

A primitive diving duck.

Pseudoseisuropsis wintu [415]

Sp. nov.

Valid

Stefanini, Gómez & Tambussi

Early Pleistocene

Miramar Formation

 Argentina

An ovenbird

Rallus nanus [416]

Nom. nov.

Valid

Alcover et al.

Holocene

 Azores

A member of Rallidae, a species of Rallus; a replacement name for Rallus minutus Alcover et al. (2015) (preoccupied).

Septencoracias [417]

Gen. et sp. nov.

Valid

Bourdon, Kristoffersen & Bonde

Eocene (Ypresian)

Fur Formation

 Denmark

A member of Coracii belonging to the family Primobucconidae. The type species is S. morsensis.

Uria onoi [418]

Sp. nov.

Valid

Watanabe et al.

Late Pleistocene

 Japan

A member of Alcidae

Wilaru prideauxi [376]

Sp. nov.

Valid

De Pietri et al.

Early Miocene

Etadunna Formation
Wipajiri Formation

 Australia

A species of Wilaru.

Pterosaurs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Allkaruen[423]

Gen. et sp. nov

Valid

Codorniú et al.

Early-Middle Jurassic

Cañadón Asfalto Formation

 Argentina

A non-pterodactyloid member of Breviquartossa. The type species is A. koi.

Aymberedactylus[424]

Gen. et sp. nov

Valid

Pêgas, Leal & Kellner

Early Cretaceous (Aptian-Albian)

Crato Formation

 Brazil

A member of Tapejarinae. The type species is A. cearensis.

Forfexopterus[425]

Gen. et sp. nov

Valid

Jiang et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Archaeopterodactyloidea. The type species is F. jeholensis.

Huaxiapterus atavismus[426]

Sp. nov

Valid

et al.

Early Cretaceous

Jiufotang Formation

 China

Pangupterus[427]

Gen. et sp. nov

Valid

et al.

Early Cretaceous

Jiufotang Formation

 China

A toothed member of Pterodactyloidea. The type species is P. liui.

Sinopterus lingyuanensis[426]

Sp. nov

Valid

et al.

Early Cretaceous

Jiufotang Formation

 China

Other reptiles

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Brasinorhynchus[437]

Gen. et sp. nov

Valid

Schultz, Langer & Montefeltro

Middle Triassic (Ladinian)

Santa Maria Formation

 Brazil

A rhynchosaur belonging to the group Stenaulorhynchinae. The type species is Brasinorhynchus mariantensis.

Colobomycter vaughni[438]

Sp. nov

Valid

MacDougall, Modesto & Reisz

Early Permian

 United States
( Oklahoma)

A member of Lanthanosuchoidea.

Euconcordia[439]

Nom. nov

Valid

Reisz, Haridy & Müller

Carboniferous (Pennsylvanian)

Calhouns Shale

 United States
( Kansas)

A member of Captorhinidae; a replacement name for Concordia Müller & Reisz (2005).

Langeronyx[440]

Gen. et comb. nov

Valid

Ezcurra, Montefeltro & Butler

Middle Triassic (Anisian)

Bromsgrove Sandstone Formation

 United Kingdom

A rhynchosaur; a new genus for "Rhynchosaurus" brodiei Benton (1990).

Ozimek[441]

Gen. et sp. nov

Valid

Dzik & Sulej

Late Triassic (probably late Carnian)

 Poland

A relative of Sharovipteryx. The type species is O. volans.

Teyujagua[442]

Gen. et sp. nov

Valid

Pinheiro et al.

Early Triassic (Induan to early Olenekian)

Sanga do Cabral Formation

 Brazil

A member of Archosauromorpha closely related to Archosauriformes. The type species is Teyujagua paradoxa.

Xinpusaurus xingyiensis[443]

Sp. nov

Valid

Li et al.

Middle Triassic (Ladinian)

 China

A thalattosaur.

Synapsids

Non-mammalian synapsids

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Abdalodon[454]

Gen. et sp. nov

Valid

Kammerer

Late Permian

Beaufort Group
(Tropidostoma Assemblage Zone)

 South Africa

An early cynodont related to Charassognathus gracilis. The type species is Abdalodon diastematicus.

Bonacynodon[455]

Gen. et sp. nov

Valid

Martinelli, Soares & Schwanke

Late Triassic (early Carnian)

Santa Maria Supersequence

 Brazil

A member of Probainognathidae. The type species is B. schultzi.

Montirictus[456]

Gen. et sp. nov

Valid

Matsuoka, Kusuhashi & Korfe

Early Cretaceous (probably Barremian or Aptian)

Kuwajima Formation

 Japan

A member of Tritylodontidae. The type species is Montirictus kuwajimaensis.

Mupashi[457]

Gen. et sp. nov

Valid

Huttenlocker & Sidor

Late Permian

Madumabisa Mudstone Formation

 Zambia

A therocephalian related to Karenites. The type species is Mupashi migrator.

Rastodon[458]

Gen. et sp. nov

Valid

Boos et al.

Permian (Guadalupian/Lopingian)

Rio do Rasto Formation

 Brazil

A dicynodont belonging to the group Bidentalia. The type species is R. procurvidens.

Santacruzgnathus[455]

Gen. et sp. nov

Valid

Martinelli, Soares & Schwanke

Late Triassic (early Carnian)

Santa Maria Supersequence

 Brazil

A member of Probainognathia, probably closely related to prozostrodontians. The type species is S. abdalai.

Sauroscaptor[459]

Gen. et sp. nov

Valid

Kammerer, Bandyopadhyay & Ray

Late Permian

Kundaram Formation

 India

A cistecephalid dicynodont. The type species is S. tharavati.

Vaughnictis[460]

Gen. et comb. nov

Valid

Brocklehurst et al.

Permian (Asselian-Sakmarian)

Cutler Group

 United States
( Colorado)

A member of Eothyrididae; a new genus for "Mycterosaurus" smithae Lewis & Vaughn (1965).

Wantulignathus[461]

Gen. et sp. nov

Valid

Whitney & Sidor

Permian (Guadalupian)

Madumabisa Mudstone Formation

 Zambia

A biarmosuchian. The type species is Wantulignathus gwembensis.

Non-eutherian mammals

Research

New taxa

Metatherians
Name Novelty Status Authors Age Unit Location Notes Images

Chaeropus baynesi[471][472]

Sp. nov

Valid

Travouillon

Late Pliocene to early Pleistocene

Moorna Formation

 Australia

A relative of the pig-footed bandicoot.

Chimeralestes[473]

Gen. et sp. nov

Valid[474]

Engelman, Anaya & Croft

Miocene

 Bolivia

A palaeothentid paucituberculate. Genus includes new species C. ambiguus.

Cookeroo[475]

Gen. et 2 sp. nov

Valid

Butler et al.

Oligocene-Miocene

Riversleigh World Heritage Area

 Australia

A member of Macropodidae. Genus includes Cookeroo bulwidarri and Cookeroo hortusensis.

Ektopodon tommosi[476]

Sp. nov

Valid

Pledge

Late Oligocene

Namba Formation

 Australia

A member of Phalangeroidea belonging to the family Ektopodontidae.

Gumardee richi[477]

Sp. nov

Valid

Travouillon et al.

Early Miocene

Riversleigh World Heritage Area

 Australia

A member of Potoroinae.

Gumardee springae[477]

Sp. nov

Valid

Travouillon et al.

Late Oligocene

Riversleigh World Heritage Area

 Australia

A member of Potoroinae.

Kutjamarcoot[478]

Gen. et sp. nov

Valid

Chamberlain et al.

Early Miocene

Wipajiri Formation

 Australia

A bandicoot. The type species is Kutjamarcoot brevirostrum.

Microleo[479]

Gen. et sp. nov

Valid

Gillespie, Archer & Hand

Early Miocene

Riversleigh World Heritage Area

 Australia

A member of Thylacoleonidae. The type species is M. attenboroughi.

Minusculodelphis modicum[480]

Sp. nov

Valid

Oliveira, Zimicz & Goin

Itaboraian

Itaboraí Quarry

 Brazil

A member of Jaskhadelphyidae, a species of Minusculodelphis.

Palaeothentes relictus[473]

Sp. nov

Valid[474]

Engelman, Anaya & Croft

Miocene

 Bolivia

A palaeothentid paucituberculate.

Palaeothentes serratus[473]

Sp. nov

Valid[474]

Engelman, Anaya & Croft

Miocene

 Bolivia

A palaeothentid paucituberculate.

Stelakoala[481]

Gen. et sp. nov

Valid

Black

Middle Miocene

Riversleigh World Heritage Area

 Australia

A member of Phascolarctidae. The type species is S. riversleighensis.

Whollydooleya[482]

Gen. et sp. nov

Valid

Archer et al.

Possibly middle or late Miocene

 Australia

A member of Dasyuridae of uncertain phylogenetic placement. The type species is W. tomnpatrichorum.

Other non-eutherian mammals
Name Novelty Status Authors Age Unit Location Notes Images

Anebodon[483]

Gen. et sp. nov

Valid

Bi et al.

Early Cretaceous

Yixian Formation

 China

A symmetrodont belonging to the family Zhangheotheriidae. The type species is A. luoi.

Cherwellia[484]

Gen. et sp. nov

Valid

Butler & Sigogneau-Russell

Middle Jurassic (late Bathonian)

 United Kingdom

A possible member of Morganucodonta. The type species is C. leei.

Culicolestes[485]

Gen. et sp. nov

Valid

Cifelli, Cohen & Davis

Late Cretaceous (Cenomanian)

Cedar Mountain Formation

 United States
( Utah)

A member of Tribosphenida of uncertain phylogenetic placement. The type species is C. kielanae.

Dakotadens pertritus[485]

Sp. nov

Valid

Cifelli, Cohen & Davis

Late Cretaceous (Cenomanian)

Cedar Mountain Formation

 United States
( Utah)

A member of Tribosphenida of uncertain phylogenetic placement.

Eotriconodon[484]

Gen. et sp. nov

Valid

Butler & Sigogneau-Russell

Middle Jurassic (late Bathonian)

 United Kingdom

A member of Triconodontidae. The type species is E. sophron.

Gobiconodon bathoniensis[484]

Sp. nov

Valid

Butler & Sigogneau-Russell

Middle Jurassic (late Bathonian)

 United Kingdom

Lactodens[486]

Gen. et sp. nov

Valid

Han & Meng

Early Cretaceous

 China

A ‘symmetrodont’ related to Spalacolestes. Genus includes new species L. sheni.

Mangasbaatar[487]

Gen. et sp. nov

Valid

Rougier et al.

Late Cretaceous (probably late Campanian)

 Mongolia

A djadochtatheriid multituberculate. The type species is M. udanii.

Morganucodon tardus[484]

Sp. nov

Valid

Butler & Sigogneau-Russell

Middle Jurassic (late Bathonian)

 United Kingdom

Phascolotherium simpsoni[484]

Sp. nov

Valid

Butler & Sigogneau-Russell

Middle Jurassic (late Bathonian)

 United Kingdom

A member of Eutriconodonta.

Stylidens[484]

Gen. et sp. nov

Valid

Butler & Sigogneau-Russell

Middle Jurassic (late Bathonian)

 United Kingdom

A possible member of Morganucodonta. The type species is S. hookeri.

Teutonodon[488]

Gen. et sp. nov

Valid

Martin et al.

Late Jurassic (Kimmeridgian)

 Germany

A plagiaulacid multituberculate. The type species is T. langenbergensis.

Theroteinus rosieriensis[489]

Sp. nov

Valid

Debuysschere

Late Triassic (Rhaetian)

 France

A member of Haramiyida belonging to the family Theroteinidae.

Eutherian mammals

Research

New taxa

Xenarthrans
Name Novelty Status Authors Age Unit Location Notes Images

Aymaratherium[537]

Gen. et sp. nov

Valid

Pujos et al.

Montehermosan

 Bolivia

A member of Nothrotheriidae. The type species is Aymaratherium jeani.

Parutaetus punaensis[538]

Sp. nov

Valid

Ciancio et al.

Middle–late Eocene

Geste Formation

 Argentina

An armadillo belonging to the subfamily Euphractinae; a species of Parutaetus.

Punatherium[538]

Gen. et sp. nov

Valid

Ciancio et al.

Middle–late Eocene

Geste Formation

 Argentina

An armadillo of uncertain phylogenetic placement. The type species is Punatherium catamarquensis.

Afrotherians
Name Novelty Status Authors Age Unit Location Notes Images

Aphanobelodon[539]

Gen. et sp. nov

Valid[540]

Wang et al.

Middle Miocene

Zhangenbao Formation

 China

A member of Amebelodontidae. The type species is A. zhaoi.

Cingulodon[541]

Gen. et sp. nov

Valid[542]

De Bast & Smith

Paleocene

 Belgium

A member of Macroscelidea belonging to the family Louisinidae. The type species is C. magioncaldai.

Eurybelodon[543]

Gen. et sp. nov

Valid

Lambert

Miocene (late Clarendonian)

Juntura Formation

 United States
( Oregon)

A gomphothere. The type species is Eurybelodon shoshanii.

Konobelodon robustus[544]

Sp. nov

Valid

Wang et al.

Late Miocene

Liushu Formation

 China

A gomphothere, a species of Konobelodon.

Prototherium ausetanum[545]

Sp. nov

Valid

Balaguer & Alba

Eocene

 Spain

A member of Dugongidae, a species of Prototherium.

Sinomammut[546]

Gen. et sp. nov

Valid

Mothé et al.

Late Miocene

Xihe-Lixian Basin

 China

A member of Mammutidae. The type species is Sinomammut tobieni.

Sinomastodon praeintermedius[547]

Sp. nov

Valid

Wang et al.

Late Miocene

Zhaotong Formation

 China

A gomphothere, a species of Sinomastodon.

Bats
Name Novelty Status Authors Age Unit Location Notes Images

Aegyptonycteris[548]

Gen. et sp. nov

Valid

Simmons, Seiffert & Gunnell

Eocene (Priabonian)

Birket Qarun Formation

 Egypt

An omnivorous bat belonging to the family Aegyptonycteridae. The type species is Aegyptonycteris knightae.

Chambinycteris[549]

Gen. et sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A bat of uncertain phylogenetic placement. The type species is C. pusilli.

Drakonycteris[549]

Gen. et sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Algeria

A bat of uncertain phylogenetic placement. The type species is D. glibzegdouensis.

Hipposideros (Pseudorhinolophus) africanum[549]

Sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A roundleaf bat.

Hipposideros (Pseudorhinolophus) amenhotepos[550]

Sp. nov

Valid

Gunnell et al.

Miocene

 Egypt

A roundleaf bat.

Khoufechia[549]

Gen. et sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A member of Nycteridae. The type species is K. gunnelli.

?Necromantis fragmentum[549]

Sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A member of Necromantidae.

?Palaeophyllophora tunisiensis[549]

Sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A member of Hipposideridae.

Premonycteris[551]

Gen. et sp. nov

In press

Hand et al.

Eocene (late Ypresian)

 France

A vesper bat. The type species is P. vesper.

Pseudovespertiliavus[549]

Gen. et sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Algeria

A sac-winged bat. The type species is P. parva.

Synemporion[552]

Gen. et sp. nov

Valid

Ziegler, Howarth & Simmons

Middle Pleistocene to late Holocene

 United States
( Hawaii)

A vesper bat. The type species is Synemporion keana.

?Vespertiliavus aenigma[549]

Sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A sac-winged bat.

Vespertiliavus kasserinensis[549]

Sp. nov

Valid

Ravel in Ravel et al.

Eocene

 Tunisia

A sac-winged bat.

Odd-toed ungulates
Name Novelty Status Authors Age Unit Location Notes Images

Cadurcodon maomingensis[553]

Sp. nov

Valid

Averianov et al.

Eocene

Youganwo Formation

 China

A member of Amynodontidae.

Hipparion philippus[554]

Sp. nov

Valid

Koufos & Vlachou

Miocene (early Turolian)

 Greece

A member of Equidae, a species of Hipparion.

Hipparion sithonis[554]

Sp. nov

Valid

Koufos & Vlachou

Miocene (early Turolian)

 Greece

A member of Equidae, a species of Hipparion.

Propalaeotherium sudrei[555]

Sp. nov

In press

Remy, Krasovec & Marandat

Eocene

 France

A member of Palaeotheriidae.

Victoriaceros hooijeri[556]

Sp. nov

Valid

Geraads et al.

Early Miocene

 Kenya

A rhinoceros, tentatively assigned to the genus Victoriaceros.

Xylotitan[557]

Gen. et sp. nov

Valid

Mihlbachler & Samuels

Eocene

Clarno Formation

 United States
( Oregon)

A member of Brontotheriidae. Genus includes new species X. cenosus.

Even-toed ungulates
Name Novelty Status Authors Age Unit Location Notes Images

Celebochoerus cagayanensis[558]

Sp. nov

Valid

Ingicco in Ingicco et al.

Middle Pleistocene

Awidon Mesa Formation

 Philippines

A member of Suidae.

Diplobunops kardoula[559]

Sp. nov

Valid

Emery, Davis & Hopkins

Eocene (Duchesnean)

 United States
( Oregon)

An agriochoerid oreodont, a species of Diplobunops.

Dorcatherium maliki[560]

Sp. nov

Valid

Kostopoulos & Sen

Miocene (Vallesian)

Ergene Formation

 Turkey

A chevrotain.

Fremdohyus[561]

Gen. et comb. nov

Valid

Prothero

Arikareean

John Day Formation

 United States
( Oregon)

A peccary. A new genus for "Thinohyus" osmonti Sinclair (1905).

Honanotherium bernori[562]

Sp. nov

Valid

Solounias & Danowitz

Late Miocene

 Iran

A member of Giraffidae, a species of Honanotherium.

Magwetherium[563]

Gen. et sp. nov

Valid

Ducrocq et al.

Eocene

Pondaung Formation

 Myanmar

A diacodexeid dichobunoid. The type species is Magwetherium burmense.

Megaloceros stavropolensis[564]

Sp. nov

Valid

Titov & Shvyreva

Early Pleistocene (late Villafranchian)

 Russia
( Stavropol Krai)

A species of Megaloceros.

Nabotherium[565]

Gen. et comb. nov

Valid

Sileem et al.

Early Oligocene

Jebel Qatrani Formation

 Egypt

An anthracothere; a new genus for "Rhagatherium" aegyptiacum Andrews (1906).

Prolibytherium fusus[566]

Sp. nov

Valid

Danowitz, Domalski & Solounias

Early Miocene

 Pakistan

A member of Ruminantia, a species of Prolibytherium.

Protovis[567]

Gen. et sp. nov

Valid

Wang, Li & Takeuchi

Pliocene

Zanda Basin

 China

A relative of the sheep. The type species is Protovis himalayensis.

Cetaceans
Name Novelty Status Authors Age Unit Location Notes Images

Acrophyseter robustus[568]

Sp. nov

Valid[569]

Lambert, Bianucci & De Muizon

Miocene (late SerravallianTortonian)

Pisco Formation

 Peru

Arktocara[570]

Gen. et sp. nov

Valid

Boersma & Pyenson

Oligocene (probably Chattian)

Poul Creek Formation

 United States
( Alaska)

A member of Platanistoidea related to Allodelphis. The type species is A. yakataga.

Awadelphis[571]

Gen. et sp. nov

Valid

Murakami

Latest Miocene

Senhata Formation

 Japan

A member of Inioidea. The type species is A. hirayamai.

Awamokoa[572]

Gen. et sp. nov

Valid[573]

Tanaka & Fordyce

Late Oligocene

Kokoamu Greensand

 New Zealand

A member of Platanistoidea. The type species is A. tokarahi.

Chavinziphius[574]

Gen. et sp. nov

Valid

Bianucci et al.

Late Miocene

Pisco Formation

 Peru

A beaked whale. The type species is C. maxillocristatus.

Chimuziphius[574]

Gen. et sp. nov

Valid

Bianucci et al.

Late Miocene

Pisco Formation

 Peru

A beaked whale. The type species is C. coloradensis.

Dagonodum[575]

Gen. et sp. nov

Valid

Ramassamy

Late Miocene

Gram Formation

 Denmark

A beaked whale. Genus includes new species D. mojnum.

Echovenator[576]

Gen. et sp. nov

Valid

Churchill et al.

Late Oligocene

Chandler Bridge Formation

 United States
( South Carolina)

A member of Xenorophidae. The type species is E. sandersi.

Fragilicetus[577]

Gen. et sp. nov

Valid

Bisconti & Bosselaers

Early Pliocene

Kattendijk Sands

 Belgium

A member of Balaenopteridae showing a mix of morphological characters that is intermediate between those of Eschrichtiidae and those of Balaenopteridae. The type species is Fragilicetus velponi.

Goedertius[578]

Gen. et. sp. nov

Valid

Kimura & Barnes

Miocene (late Aquitanian)

Nye Formation

 United States ( Oregon)

A member of Allodelphinidae. The type species is Goedertius oregonensis.

Incakujira[579]

Gen. et sp. nov

Valid

Marx & Kohno

Miocene

Pisco Formation

 Peru

A rorqual. The type species is I. anillodefuego.

Mammalodon hakataramea[580]

Sp. nov

Valid

Fordyce & Marx

Oligocene (Duntroonian, 27 Ma)

Kokoamu Greensand

 New Zealand

A species of Mammalodon, a member of Mammalodontidae.

Matapanui[581][582]

Gen. et sp. nov

Valid[583]

Boessenecker & Fordyce

Oligocene (latest Whaingaroan, 28.1–27.3 Ma)

Kokoamu Greensand

 New Zealand

A member of Eomysticetidae. The type species is M. waihao.

Mesoplodon posti[584]

Sp. nov

Valid

Lambert & Louwye

Pliocene (Zanclean)

 Belgium

A mesoplodont whale.

Metopocetus hunteri[585]

Sp. nov

Valid

Marx, Bosselaers & Louwye

Miocene (late Tortonian)

Breda Formation

 Netherlands

A member of Cetotheriidae, a species of Metopocetus.

Mithridatocetus[586]

Gen. et sp. et comb. nov

Valid

Gol'din & Startsev

Miocene (Tortonian)

Chersonian Formation

 Ukraine Russia

A member of Cetotheriidae. Genus includes new species M. eichwaldi, as well as "Kurdalagonus" adygeicus Tarasenko & Lopatin (2012) and "Cetotherium" mayeri (a nomen dubium).

Ninjadelphis[578]

Gen. et. sp. nov

Valid

Kimura & Barnes

Miocene (late Burdigalian)

Hiramatsu Formation

 Japan

A member of Allodelphinidae. The type species is Ninjadelphis ujiharai.

Rayanistes[587]

Gen. et sp. nov

Valid

Bebej et al.

Eocene (Lutetian)

Midawara Formation

 Egypt

A member of Remingtonocetidae. The type species is Rayanistes afer.

Sitsqwayk[588]

Gen. et sp. nov

Valid

Peredo & Uhen

Late Oligocene

Pysht Formation

 United States
( Washington)

A basal member of Chaeomysticeti. The type species is S. cornishorum.

Whakakai[589]

Gen. et sp. nov

Valid

Tsai & Fordyce

Oligocene (Chattian)

Kokoamu Greensand

 New Zealand

A baleen whale. The type species is Whakakai waipata.

Zarhinocetus donnamatsonae[578]

Sp. nov

Valid

Kimura & Barnes

Miocene (late Burdigalian-early Langhian)

Astoria Formation

 United States ( Washington)

A member of Allodelphinidae.

Carnivorans
Name Novelty Status Authors Age Unit Location Notes Images

Actiocyon parverratis[590]

Sp. nov

Valid

Smith, Czaplewski & Cifelli

Miocene (Barstovian)

Monarch Mill Formation

 United States
( Nevada)

A simocyonine ailurid (a relative of the red panda), a species of Actiocyon.

Alagtsavbaatar[591]

Gen. et comb. nov

Valid

Egi et al.

Late Eocene

Ergilin Dzo Formation

 Mongolia

A member of Feliformia. The type species is "Stenoplesictis" indigenus Dashzeveg (1996).

Angelarctocyon[592]

Gen. et comb. nov

Valid

Tomiya & Tseng

Eocene (Duchesnean)

Chambers Tuff

 United States
( Texas)

A bear dog. The type species is "Miacis" australis Gustafson (1986).

Brevimalictis[590]

Gen. et sp. nov

Valid

Smith, Czaplewski & Cifelli

Miocene (Barstovian)

Monarch Mill Formation

 United States
( Nevada)

A member of Mustelidae of uncertain phylogenetic placement. The type species is Brevimalictis chikasha.

Cynarctus wangi[593]

Sp. nov

Valid

Jasinski & Wallace

Middle Miocene

Choptank Formation

 United States
( Maryland)

A member of Canidae belonging to the subfamily Borophaginae, a species of Cynarctus.

Enhydrictis hoffstetteri[594]

Sp. nov

In press

Geraads

Pleistocene

 Algeria

An otter-like member of the Mustelidae.

Gustafsonia[592]

Gen. et comb. nov

Valid

Tomiya & Tseng

Eocene (Chadronian)

Chambers Tuff

 United States
( Texas)

A bear dog. The type species is "Miacis" cognitus Gustafson (1986).

Lontra weiri[595]

Sp. nov

Valid

Prassack

Pliocene

Hagerman Fossil Beds

 United States
( Idaho)

An otter, a species of Lontra.

Lutraeximia[596]

Gen. et sp. nov

Valid

Cherin et al.

Early Pleistocene (late Villafranchian)

 Italy

An otter. The type species is Lutraeximia umbra.

Maofelis[597]

Gen. et sp. nov

Valid

Averianov et al.

Middle-late Eocene

Youganwo Formation

 China

A member of Nimravidae. The type species is Maofelis cantonensis.

Megantereon microta[598]

Sp. nov

In press

Zhu et al.

Early Pleistocene

 China

A machairodontine felid, a species of Megantereon.

Negodiaetictis[590]

Gen. et sp. nov

Valid

Smith, Czaplewski & Cifelli

Miocene (Barstovian)

Monarch Mill Formation

 United States
( Nevada)

A member of Mustelidae of uncertain phylogenetic placement. The type species is Negodiaetictis rugatrulleum.

Watay[599]

Gen. et sp. nov

Valid

McLaughlin, Hopkins & Schmitz

Hemingfordian

Mascall Formation

 United States
( Oregon)

A member of Mustelidae. The type species is W. tabutsigwii.

Rodents
Name Novelty Status Authors Age Unit Location Notes Images

Apatodemus[600]

Gen. et sp. nov

Valid

Savorelli, Colombero & Masini

Miocene

 Italy

A member of Muridae. The type species is A. degiulii.

Argouburus[601]

Gen. et sp. nov

Valid[602]

Marivaux et al.

Early Oligocene

Samlat Formation

Western Sahara

An anomalure. The type species is A. minutus.

Birkamys[603]

Gen. et sp. nov

Valid

Sallam & Seiffert

Eocene (late Priabonian)

Jebel Qatrani Formation

 Egypt

A member of Phiomorpha of uncertain phylogenetic placement. The type species is Birkamys korai.

Brachyscirtetes tomidai[604]

Sp. nov

Valid

Li

Late Miocene

 China

A member of Dipodidae, a species of Brachyscirtetes.

Chambiramys[605]

Gen. et 2 sp. nov

Valid

Boivin et al.

Late Oligocene

Chambira Formation

 Peru

A caviomorph rodent of uncertain phylogenetic placement. The type species is C. sylvaticus; genus also includes C. shipiborum.

Chukimys[606]

Gen. et sp. nov

Valid

Barbiere et al.

Late Pliocene

Brochero Formation

 Argentina

A member of Sigmodontinae. The type species is C. favaloroi.

Cricetodon fikreti[607]

Sp. nov

Valid

Çinar Durgut & Ünay

Early Miocene

 Turkey

A member of Cricetodontini.

Cricetodon magnesiensis[607]

Sp. nov

Valid

Çinar Durgut & Ünay

Early Miocene

 Turkey

A member of Cricetodontini.

Cricetodon trallesensis[607]

Sp. nov

Valid

Çinar Durgut & Ünay

Early Miocene

 Turkey

A member of Cricetodontini.

Cricetodon yapintiensis[607]

Sp. nov

Valid

Çinar Durgut & Ünay

Early Miocene

 Turkey

A member of Cricetodontini.

Dakhlamys[601]

Gen. et sp. nov

Valid[602]

Marivaux et al.

Early Oligocene

Samlat Formation

Western Sahara

A possible member of Zegdoumyidae. The type species is D. ultimus.

Elfomys catalaunicus[608]

Sp. nov

Valid

Bonilla-Salomón et al.

Eocene

 Spain

A member of Theridomyidae, a species of Elfomys.

Elymys ? emryi[609]

Sp. nov

Valid

Kelly & Murphey

Eocene (early Uintan)

Turtle Bluff Member, Bridger Formation

 United States
( Wyoming)

A possible member of Dipodidae.

Eucricetodon wangae[610]

Sp. nov

Valid

Li, Meng & Wang

Late Eocene

 China

A member of Cricetidae, a species of Eucricetodon.

Geomys tyrioni[611]

Sp. nov

Valid

Martin

Early Pleistocene

Meade Basin

 United States
( Kansas)

A pocket gopher, a species of Geomys.

Gliruloides[612]

Gen. et sp. et comb. nov

Valid

Wu et al.

Late Oligocene to early Miocene

Junggar Basin

 China
 Turkey

A dormouse related to the forest dormouse. The type species is Gliruloides zhoui; genus also includes "Vasseuromys" duplex Ünay (1994).

Irtyshogaulus[613]

Gen. et 2 sp. nov

Valid

Lu et al.

Early Miocene

Junggar Basin

 China

A member of Mylagaulidae belonging to the subfamily Promylagaulinae. The type species is I. minor; genus also includes I. major.

Lagostomus telenkechanum[614]

Sp. nov

Valid

Rasia & Candela

Late Miocene

Arroyo Chasicó Formation

 Argentina

A species of Lagostomus.

Loretomys[605]

Gen. et sp. nov

Valid

Boivin et al.

Late Oligocene

Chambira Formation

 Peru

A caviomorph rodent belonging to the superfamily Octodontoidea. The type species is L. minutus.

Megacricetodon hellenicus[615]

Sp. nov

Valid

Oliver & Peláez-Campomanes

Early Miocene

 Greece

A cricetid rodent, a species of Megacricetodon.

Mioheteromys subterior[616]

Sp. nov

Valid

Korth & Evander

Miocene (early Barstovian)

 United States
( Nebraska)

A heteromyid rodent, a species of Mioheteromys.

Mimomys chandolensis[617]

Sp. nov

Valid

Tiunov, Golenishchev & Voyta

Late Pleistocene

 Russia
( Primorsky Krai)

An arvicoline cricetid, a species of Mimomys.

Minocastor[618]

Gen. et sp. nov

Valid

Mörs, Tomida & Kalthoff

Early Miocene

Nakamura Formation

 Japan

A member of Castoridae. The type species is Minocastor godai.

Mubhammys[603]

Gen. et sp. nov

Valid

Sallam & Seiffert

Eocene (late Priabonian)

Jebel Qatrani Formation

 Egypt

A member of Phiomorpha of uncertain phylogenetic placement. The type species is Mubhammys vadumensis.

Myocricetodon tomidai[619]

Sp. nov

Valid

Lindsay & Flynn

Early Miocene

Chitarwata Formation
Vihowa Formation

 Pakistan

A member of Cricetidae, a species of Myocricetodon.

Mystemys[620]

Gen. et sp. nov

Valid

Savorelli & Masini

Miocene

 Italy

A member of Cricetidae. The type species is M. giganteus.

Neochoerus occidentalis[621]

Sp. nov.

In press

Carranza y Castañeda

Late Blancan-Irvingtonian

 Mexico

Nonanomalurus parvus[601]

Sp. nov

Valid[602]

Marivaux et al.

Early Oligocene

Samlat Formation

Western Sahara

A member of Anomaluroidea.

?Notoparamys blochi[622]

Sp. nov

Valid

Gunnell, Zonneveld & Bartels

Wasatchian

Wasatch Formation

 United States
( Wyoming)

A member of Ischyromyidae belonging to the subfamily Paramyinae, tentatively assigned to the genus Notoparamys.

Oromys[601]

Gen. et sp. nov

Valid[602]

Marivaux et al.

Early Oligocene

Samlat Formation

Western Sahara

A member of Zenkerellinae. The type species is O. zenkerellinopsis.

Paciculus walshi[623]

Sp. nov

Valid

Lindsay et al.

Oligocene

Otay Formation

 United States
( California)

A member of Cricetidae, a species of Paciculus.

Palaeosteiromys[605]

Gen. et sp. nov

Valid

Boivin et al.

Late Oligocene

Chambira Formation

 Peru

A New World porcupine. The type species is P. amazonensis.

Pappocricetodon siziwangqiensis[610]

Sp. nov

Valid

Li, Meng & Wang

Late Eocene

 China

A member of Cricetidae, a species of Pappocricetodon.

Paradelomys santjaumensis[608]

Sp. nov

Valid

Bonilla-Salomón et al.

Eocene

 Spain

A member of Theridomyidae, a species of Paradelomys.

Paranomalurus riodeoroensis[601]

Sp. nov

Valid[602]

Marivaux et al.

Early Oligocene

Samlat Formation

Western Sahara

An anomalure.

Plesiosciurus zhengi[624]

Sp. nov

Valid

Qiu & Jin

Cenozoic (probably Miocene)

 China

A member of Sciurini.

Plesiosteiromys[605]

Gen. et sp. nov

Valid

Boivin et al.

Late Oligocene

Chambira Formation

 Peru

A caviomorph rodent, possiby a New World porcupine. The type species is P. newelli.

Postcopemys chapalensis[625]

Sp. nov

Valid

Rincón et al.

Early Pliocene

Chapala Formation

 Mexico

A member of Cricetidae.

Potamarchus adamiae[626]

Sp. nov

Valid

Kerber et al.

Late Miocene

Solimões Formation

 Brazil

A potamarchine dinomyid, a species of Potamarchus.

Potwarmus mahmoodi[619]

Sp. nov

Valid

Lindsay & Flynn

Early Miocene

Vihowa Formation

 Pakistan

A member of Cricetidae, a species of Potwarmus.

Primus cheemai[619]

Sp. nov

Valid

Lindsay & Flynn

Late Oligocene to Early Miocene

Chitarwata Formation
Vihowa Formation

 Pakistan

A member of Cricetidae, a species of Primus.

Proansomys badamae[627]

Sp. nov

Valid

Maridet et al.

Late Oligocene

Hsanda Gol Formation

 Mongolia

A member of Aplodontiidae.

Pseudopotamarchus[626]

Gen. et sp. nov

Valid

Kerber et al.

Late Miocene

Solimões Formation

 Brazil

A potamarchine dinomyid. The type species is Pseudopotamarchus villanuevai.

Pseudoratufa[624]

Gen. et sp. nov

Valid

Qiu & Jin

Cenozoic (probably Miocene)

 China

A relative of the oriental giant squirrels. The type species is P. wanensis.

Sayimys negevensis[628]

Sp. nov

Valid

López-Antoñanzas et al.

Early Miocene

Middle Hatzeva Formation

 Israel

A gundi, a species of Sayimys.

Scleromys praecursor[605]

Sp. nov

Valid

Boivin et al.

Late Oligocene

Chambira Formation

 Peru

A caviomorph rodent belonging to the family Dinomyidae.

Ucayalimys[605]

Gen. et sp. nov

Valid

Boivin et al.

Late Oligocene

Chambira Formation

 Peru

A caviomorph rodent, possibly a member of the superfamily Chinchilloidea. The type species is U. crassidens.

Ullumys[629]

Gen. et sp. et comb. nov

Valid

Olivares et al.

Miocene (Huayquerian)

Las Tapias Formation

 Argentina

A member of Echimyidae. Genus includes new species U. pattoni, as well as "Eumysops" intermedius Rovereto.

Primates
Name Novelty Status Authors Age Unit Location Notes Images

Agerinia smithorum[630]

Sp. nov

Valid

Femenias-Gual et al.

Early Eocene

 Spain

Apidium zuetina[631]

Sp. nov

Valid

Beard et al.

Early Oligocene

 Libya

A species of Apidium.

Bahinia banyueae[632]

Sp. nov

Valid

Ni et al.

Early Oligocene

Caijiachong Formation

 China

A member of Eosimiidae, a species of Bahinia.

Canaanimico[633]

Gen. et sp. nov

Valid

Marivaux et al.

Late Oligocene

Chambira Formation

 Peru

A New World monkey related to Soriacebus. The type species is C. amazonensis.

Cantius lohseorum[634]

Sp. nov

Valid

Robinson

Early Eocene

Powder River Basin

 United States
( Wyoming)

A member of Notharctidae.

Gatanthropus[632]

Gen. et sp. nov

Valid

Ni et al.

Early Oligocene

Caijiachong Formation

 China

A relative of Ekgmowechashala. The type species is Gatanthropus micros.

Laomaki[632]

Gen. et sp. nov

Valid

Ni et al.

Early Oligocene

Caijiachong Formation

 China

A member of Adapiformes belonging to the family Sivaladapidae. The type species is Laomaki yunnanensis.

Megaceralemur[634]

Gen. et comb. et sp. nov

Valid

Robinson

Eocene

 United States
( Wyoming)

A member of Notharctidae. A new genus for "Pelycodus" trigonodus Matthew (1915); genus also includes new species M. matthewi.

Oligotarsius[632]

Gen. et sp. nov

Valid

Ni et al.

Early Oligocene

Caijiachong Formation

 China

A tarsier-like primate. The type species is Oligotarsius rarus.

Panamacebus[635]

Gen. et sp. nov

Valid

Bloch et al.

Early Miocene

Las Cascadas Formation

 Panama

A New World monkey, probably a member of Cebidae. The type species is Panamacebus transitus.

Pinolophus[634]

Gen. et sp. nov

Valid

Robinson

Eocene

 United States
( Wyoming)

A member of Notharctidae. The type species P. meikei.

Semnopithecus gwebinensis[636]

Sp. nov

Valid

Takai et al.

Late Pliocene

 Myanmar

A gray langur.

Yunnanadapis[632]

Gen. et 2 sp. nov

Valid

Ni et al.

Early Oligocene

Caijiachong Formation

 China

A member of Adapiformes belonging to the family Sivaladapidae. The type species is Yunnanadapis folivorus; genus also includes Yunnanadapis imperator.

Other eutherians
Name Novelty Status Authors Age Unit Location Notes Images

Akhnatenavus nefertiticyon[637]

Sp. nov

Valid

Borths, Holroyd & Seiffert

Eocene (latest Priabonian)

Jebel Qatrani Formation

 Egypt

A hyainailourine hyaenodont.

Amphilagus tomidai[638]

Sp. nov

Valid

Erbajeva, Angelone & Alexeeva

Miocene

 Russia
( Irkutsk Oblast)

A lagomorph, a species of Amphilagus.

Antesorex wilsoni[616]

Sp. nov

Valid

Korth & Evander

Miocene (early Barstovian)

 United States
( Nebraska)

A shrew, a species of Antesorex.

Belgoryctes[541]

Gen. et sp. nov

Valid[542]

De Bast & Smith

Paleocene

 Belgium

A member of Palaeoryctidae. The type species is B. thaleri.

Brychotherium[637]

Gen. et sp. nov

Valid

Borths, Holroyd & Seiffert

Eocene (latest Priabonian)

Jebel Qatrani Formation

 Egypt

A teratodontine hyaenodont. The type species is B. ephalmos.

Cambaytherium gracilis[150]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

Cambay Shale Formation

 India

A perissodactyl-like ungulate.

Eoictops[622]

Gen. et sp. nov

Valid

Gunnell, Zonneveld & Bartels

Wasatchian

Wasatch Formation

 United States
( Wyoming)

A member of Leptictida belonging to the family Leptictidae. The type species is E. novaceki.

Eurolestes[541]

Gen. et sp. nov

Valid[542]

De Bast & Smith

Paleocene

 Belgium

A member of Pentacodontidae. The type species is E. dupuisi.

Gomphos progressus[639]

Sp. nov

Valid

Li, Wang & Fostowicz-Frelik

Eocene (Irdinmanhan)

Ulan Shireh Formation

 China

A member of Glires belonging to the family Mimotonidae.

Hegetotherium cerdasensis[640]

Sp. nov

Valid

Croft et al.

Miocene (Langhian)

 Bolivia

A notoungulate, a species of Hegetotherium.

Indoesthonyx[150]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

Cambay Shale Formation

 India

A member of Tillodontia. The type species is I. suratensis.

Lanthanotherium observatum[616]

Sp. nov

Valid

Korth & Evander

Miocene (early Barstovian)

 United States
( Nebraska)

A member of Erinaceidae, a species of Lanthanotherium.

Mina[641][642]

Gen. et sp. nov

Valid

Li et al.

Middle Paleocene

Wanghudun Formation

 China

A basal member of Glires. The type species is Mina hui.

Ounalashkastylus[643]

Gen. et sp. nov

Valid

Chiba et al.

Miocene

 United States
( Alaska)

A desmostylian. The type species is Ounalashkastylus tomidai.

Palaeictops altimontis[644]

Sp. nov

Valid

Velazco & Novacek

Eocene (Uintan)

Tepee Trail Formation

 United States
( Wyoming)

A member of Leptictida belonging to the family Leptictidae.

Palaeictops robustus[644]

Sp. nov

Valid

Velazco & Novacek

Eocene (Uintan)

Uinta Formation

 United States
( Utah)

A member of Leptictida belonging to the family Leptictidae.

Palaeosinopa lacus[622]

Sp. nov

Valid

Gunnell, Zonneveld & Bartels

Wasatchian

Wasatch Formation

 United States
( Wyoming)

A member of Pantolestidae.

Pternoconius bondi[645]

Sp. nov

Valid

Cheme Arriaga, Dozo & Gelfo

Miocene (Colhuehuapian)

Sarmiento Formation

 Argentina

A member of Litopterna belonging to the family Macraucheniidae and the subfamily Cramaucheniinae.

Ptilocercus kylin[646]

Sp. nov

Valid

Li & Ni

Earliest Oligocene

Lijiawa Mammalian Fossil locality

 China

A treeshrew related to the pen-tailed treeshrew.

Purgatorius pinecreeensis[647]

Sp. nov

Valid

Scott, Fox & Redman

Paleocene (Puercan)

Ravenscrag Formation

 Canada
( Saskatchewan)

Quadratodon[541]

Gen. et sp. nov

Valid[542]

De Bast & Smith

Paleocene

 Belgium

A member of Erinaceomorpha. The type species is Q. sigei.

Rzebikia[648]

Gen. et comb. nov

Valid

Sansalone, Kotsakis & Piras

Pliocene to Pleistocene (Villanyian)

 Bulgaria
 Poland

A relative of the American shrew mole. A new genus for "Neurotrichus" polonicus Skoczeń (1980) and "Neurotrichus" skoczeni Zijlstra (2010).

Sanshuilophus[649]

Gen. et sp. nov

Valid

Mao et al.

Early Eocene

Huayong Formation

 China

A member of Phenacolophidae (a group of archaic ungulate mammals of uncertain phylogenetic placement). The type species is Sanshuilophus zhaoi.

Scalopoides hutchisoni[616]

Sp. nov

Valid

Korth & Evander

Miocene (early Barstovian)

 United States
( Nebraska)

A member of Talpidae, a species of Scalopoides.

Trachytherus ramirezi[650]

Sp. nov

Valid

Shockey, Billet & Salas-Gismondi

Oligocene (Deseadan)

 Peru

A mesotheriid notoungulate, a species of Trachytherus.

Vassacyon prieuri[651]

Sp. nov

Valid

Solé et al.

Latest Paleocene

 France

A basal member of Carnivoraformes, a species of Vassacyon.

Zofialestes[652]

Gen. et sp. nov

Valid

Fostowicz-Frelik

Late Cretaceous (?late Campanian)

Barun Goyot Formation

 Mongolia

A relative of Zalambdalestes. The type species is Z. longidens.

Other animals

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Acoelia norica[660]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

 Turkey

A demosponge belonging to the group Agelasida.

Annulitubus[661]

Gen. et sp. nov

Valid

Vinn et al.

Devonian (late Emsian)

Ponta Grossa Formation

 Brazil

An annelid, possibly a polychaete. The type species is Annulitubus mutveii.

Antennipatus[662]

Gen. et sp. nov

Valid

Garwood, Edgecombe & Giribet in Garwood et al.

Carboniferous (Stephanian)

Montceau-les-Mines Lagerstätte

 France

A velvet worm. The type species is A. montceauensis.

Archiasterella dhiraji[663]

Sp. nov

Valid

Gilbert, Hughes & Myrow

Cambrian

Parahio Formation

 India

Bistella[664]

Gen. et sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A heteractinid sponge belonging to the group Heteractinellida and the family Heterostellidae. The type species is B. inexplicabila.

Bohemograptus praecox[665]

Sp. nov

Valid

Štorch et al.

Silurian

 Czech Republic

A graptolite.

Bolidium bertii[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Bolidium.

Burejospermum punctatum[668]

Sp. nov

Valid

McLoughlin et al.

Early Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A member of Clitellata of uncertain phylogenetic placement, described on the basis of fossilized cocoons; a species of Burejospermum.

Burejospermum seymourense[668]

Sp. nov

Valid

McLoughlin et al.

Early Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A member of Clitellata of uncertain phylogenetic placement, described on the basis of fossilized cocoons; a species of Burejospermum.

Caelispongia[660]

Gen. et sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Stellispongiellidae. The type species is C. topukensis.

Camerospongia tuberculata[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Camerospongia.

Camerospongia visentinae[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Camerospongia.

Capsulocyathus petri[669]

Sp. nov

Valid

Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Cryptoporocyathidae.

Cavispongia scarpai[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Cavispongia.

Conotheca hensoni[670]

Sp. nov

Valid

Peel et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Hyolitha belonging to the group Orthothecida.

Constellatispongia[671]

Gen. et sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge. The type species is C. canismajorii.

Corallistes multiosculata[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Corallistes.

Cornulites vilcae[672]

Sp. nov

Valid

Vinn & Gutiérez-Marco

Late Ordovician (probably late Sandbian)

Probably Calapuja Formation

 Peru

A member of Cornulitida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Cornulites.

Cornulites zatoni[672]

Sp. nov

Valid

Vinn & Gutiérez-Marco

Late Ordovician (Sandbian)

Calapuja Formation

 Peru

A member of Cornulitida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Cornulites.

Coronispongia[666]

Gen. et sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge. The type species is Coronispongia confossa.

Crassicoactum[671]

Gen. et sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge. The type species is C. cucumis.

Daharella triassica[660]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Auriculospongiidae.

Eolipastrotethya[673]

Gen. et sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Bubarida. The type species is E. picketti.

Esakovella[664]

Gen. et sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A tommotiid belonging to the family Kelanellidae. The type species is E. grigorievae.

Fallocyathus accomodatus[669]

Sp. nov

Valid

Sundukov in Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Fallocyathidae.

Fallocyathus apheles[669]

Sp. nov

Valid

Sundukov in Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Fallocyathidae.

Geodia hopetouni[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida, a species of Geodia.

Hadimopanella incubo[675]

Sp. nov

Valid[676]

Streng, Ebbestad & Berg-Madsen

Cambrian

 Sweden

A palaeoscolecid worm.

Hadimopanella oelandiana[675]

Sp. nov

Valid[676]

Streng, Ebbestad & Berg-Madsen

Cambrian

 Sweden

A palaeoscolecid worm.

Hamptonia limatula[671]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Hexactinella clampensis[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Hexactinella.

?Hyalostelia spinula[664]

Sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A hexactinellid sponge belonging to the group Hexasterophora and the family Hyalostellidae.

Lenica perversa[671]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Lyrarapax trilobus[677]

Sp. nov

Valid

Cong et al.

Early Cambrian

 China

Milyasa[660]

Gen. et 2 sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Dereköy Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Preperonidellidae. The type species is M. polysiphonata; genus also includes M. askomorpha.

Monocrepidium pauli[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Bubarida.

Monocrepidium unispiculatum[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Bubarida.

Nganki namak[678]

Sp. nov

Valid

Kruse & Hughes

Cambrian

Khussak Formation

 Pakistan

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Nganki.

Obscurospongia[660]

Gen. et sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

An inozoid or chaetetid sponge. The type species is O. chaetetiformis

Ottoia guizhouensis[679]

Sp. nov

Valid

Yang, Zhao & Zhang

Cambrian

Kaili Formation

 China

A stem-group priapulid, a species of Ottoia.

Pachastrella australis[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida.

Pachastrella intermedia[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida.

Palaeoconchus wilsoni[680]

Sp. nov

Valid

Zatoń, Vinn & Toom

Silurian (Ludlow)

Kuressaare Formation

 Estonia
 Sweden

A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs).

Palaeotubus[681]

Gen. et sp. nov

Valid

Sanfilippo et al.

Permian

Pietra di Salomone limestone

 Italy

A polychaete, possibly a relative of serpulids. The type species is P. sosiensis.

Pegmatothylakos[668]

Gen. et sp. nov

Valid

McLoughlin et al.

Early Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A member of Clitellata of uncertain phylogenetic placement, described on the basis of fossilized cocoons. The type species is Pegmatothylakos manumii.

Phycopsis arbusculum[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Axinellida.

Preperonidella asymmetrica[660]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Preperonidellidae.

Pseudosardospongia[664]

Gen. et 2 sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A calcareous sponge belonging to the family Polyactinellidae. The type species is P. plana; genus also includes P. miriculata.

Rajatubulus[682]

Gen. et comb. nov

Valid

Yang et al.

Early Cambrian

 Kazakhstan

A member of Cloudinidae; a new genus for "Pseudorthotheca" costata Mambetov in Missarzhevsky & Mambetov (1981).

Rankenella zhangxianensis[683]

Sp. nov

Valid

Lee, Woo & Lee

Cambrian

Zhangxia Formation

 China

A sponge belonging to the family Anthaspidellidae.

Ratcliffespongia freuchenensis[671]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Rigonia[666]

Gen. et sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge. The type species is Rigonia plicata.

Rotundocyathus indistinctus[669]

Sp. nov

Valid

Sundukov in Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Ajacicyathidae.

Rugatotheca daibuica[682]

Sp. nov

Valid

Yang et al.

Early Cambrian

Daibu Member of the Xiaotan section

 China

A small shelly fossil of uncertain phylogenetic placement.

Saccoglossus testa[684]

Sp. nov

Valid

Cameron

Carboniferous (Pennsylvanian)

Mazon Creek fossil beds

 United States
( Illinois)

An acorn worm belonging to the family Harrimaniidae, a species of Saccoglossus.

Saetaspongia procera[671]

Sp. nov

Valid

Botting & Peel

Early Cambrian

Buen Formation

 Greenland

A sponge.

Sanxiascolex[685]

Gen. et sp. nov

Valid

Yang & Zhang

Cambrian

Shipai Formation

 China

A palaeoscolecid. The type species is P. papillogyrus.

Sardospongia gigantea[664]

Sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A calcareous sponge belonging to the family Polyactinellidae.

Sardospongia triplexa[664]

Sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A calcareous sponge belonging to the family Polyactinellidae.

Sarmentofascis zamparelliae[686]

Sp. nov

Valid

Schlagintweit, Frijia & Parente

Late Cretaceous (early Campanian)

 Italy

A sponge, a species of Sarmentofascis.

Sibirecyathus onkhoydokh[669]

Sp. nov

Valid

Sundukov & Zhuravlev

Cambrian

 Russia

A member of Archaeocyatha belonging to the family Ajacicyathidae.

Stauractinella eocenica[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Stauractinella.

Talacastospongia[687]

Gen. et sp. nov

Valid

Carrera & Rustán

Devonian (Lochkovian)

Talacasto Formation

 Argentina

A hexactinellid sponge. The type species is Talacastospongia minima.

Tanchajella[664]

Gen. et sp. nov

Valid

Fedorov in Fedorov, Parkhaev & Demidenko

Cambrian

 Russia

A hexactinellid sponge belonging to the group Amphidiscophora and the family Hyalonematidae. The type species is T. aculeata.

Taurispongia[660]

Gen. et 8 sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Dereköy Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Stellispongiellidae. The type species is T. oligocanalis; genus also includes new species T. polycanalis, T. tenuis, T. lamellicanalis, T. fascifera, T. siderifera, T. lamellata and T. polyforma.

Tenuipariespongia taurica[660]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Kasımlar Basin

 Turkey

A demosponge belonging to the group Agelasida.

Tritonychus [688]

Gen. et sp. nov.

Valid

Zhang et al.

Early Cambrian

 China

A lobopodian. The type species is T. phanerosarkus.

Tulenicornus? frykmani[670]

Sp. nov

Valid

Peel et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Hyolitha belonging to the group Hyolithida.

Toulminia italica[666]

Sp. nov

Valid[667]

Frisone, Pisera & Preto

Eocene (early Lutetian)

 Italy

A sponge, a species of Toulminia.

Triptolemma solida[673]

Sp. nov

Valid[674]

Łukowiak & Pisera

Late Eocene

Pallinup Formation

 Australia

A heteroscleromorph demosponge belonging to the group Tetractinellida.

Tyanada[660]

Gen. et comb. et sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian-Rhaetian)

 Australia
 Austria
 Iran
 Turkey

A demosponge belonging to the group Agelasida and the family Virgulidae. The type species is "Grossotubenella" variabilis Senowbari-Daryan (2005); genus also includes new species T. irregularis.

Wildspongia? krystyni[660]

Sp. nov

Valid

Senowbari-Daryan & Link

Late Triassic (Norian)

Dereköy Basin

 Turkey

A demosponge belonging to the group Agelasida and the family Stellispongiellidae.

Wronascolex geyiensis[689]

Sp. nov

Valid

Peng et al.

Cambrian

 China

A palaeoscolecid worm, a species of Wronascolex.

Wronascolex? johanssoni[675]

Sp. nov

Valid[676]

Streng, Ebbestad & Berg-Madsen

Cambrian

 Sweden

A palaeoscolecid worm.

Wronascolex yichangensis[690]

Sp. nov

Valid

Yang & Zhang

Cambrian

 China

A palaeoscolecid worm, a species of Wronascolex.

Other organisms

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Accordiella? tarburensis[696]

Sp. nov

Valid

Schlagintweit & Rashidi

Late Cretaceous (Maastrichtian)

Tarbur Formation

 Iran

A foraminifer belonging to the group Textulariida and the family Chrysalidinidae.

Acuasiphonoria[697]

Gen. et sp. nov

Valid

Liu et al.

Ordovician

Tarim Basin

 China

A member of Cyanobacteria. The type species is Acuasiphonoria ordovica.

Adnatosphaeridium ivoriense[698]

Sp. nov

Valid

Awad & Oboh-Ikuenobe

Late Paleocene and early Eocene

Gulf of Guinea (Côte D'Ivoire-Ghana Transform Margin)
 Nigeria

A dinoflagellate belonging to the group Gonyaulacales.

Agathammina vachardi[699]

Sp. nov

Valid

Zhang in Zhang et al.

Permian

Xiala Formation

 China

A foraminifer belonging to the group Miliolata and the family Cornuspiridae.

Amsassia koreanensis[700]

Sp. nov

Valid

Lee et al.

Middle and Late Ordovician (Darriwilian-Katian)

Duwibong Formation
Xiazhen Formation

 China
 South Korea

An organism of uncertain phylogenetic placement, probably a calcareous alga.

Anchisolenopora[701]

Gen. et comb. nov

Valid

Vachard et al.

Ordovician-Carboniferous

 China
 France
 Norway
 Sweden

A red alga, possibly related to the coralline algae. The type species is "Hedstroemia" serrana Vachard & Aretz (2004); genus also includes "Hedstroemia" nidarosiensis Høeg, 1932 emend. Roux, 1985, "Hedstroemia" koninckoporoides Vachard, 1988 and "Pseudosolenopora" owodenkoi sensu Mamet, 2002 non Chanton-Güvenç, 1972.

Annelaurea[702]

Gen. et sp. nov

Valid

Harper et al.

Carboniferous (late Viséan)

 France

An oomycete. The type species is Annelaurea excornis.

Aulacoseira helianthus[703]

Sp. nov

Valid

Mohan & Stone in Mohan, Stone & Campisano

Pliocene

Hadar Formation

 Ethiopia

A diatom belonging to the group Aulacoseirales and the family Aulacoseiraceae.

Aulacoseira jewsonii[703]

Sp. nov

Valid

Mohan & Stone in Mohan, Stone & Campisano

Pliocene

Hadar Formation

 Ethiopia

A diatom belonging to the group Aulacoseirales and the family Aulacoseiraceae.

Barattolites andhuri[704]

Sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Eocene

Dammam Formation

 Oman
 Yemen

A foraminifer belonging to the group Globothalamea and the family Orbitolinidae.

Berkutaphycus[705]

Gen. et sp. nov

Valid

Schopf, Sergeev & Kudryavtsev

Early Cambrian

Kyrshabakta Formation

 Kazakhstan

An organism of uncertain phylogenetic placement; might be related to hormogonian cyanobacteria or to eukaryotic green or chrysophyte algae. The type species is Berkutaphycus elongatus.

Bianchina[706]

Gen. et sp. nov

Valid

Schiøler

Cretaceous (late Aptian–middle Cenomanian)

East Coast Basin

 New Zealand

A dinoflagellate. The type species is Bianchina hieroglyphica.

Braarudosphaera wendleriae[707]

Sp. nov

Valid

Lees & Bown

Late Cretaceous (Turonian)

 Tanzania

A microfossil.

Broccoliforma[708]

Gen. et sp. nov

Valid

Mason & Narbonne

Ediacaran

 Canada
( Newfoundland and Labrador)

A flabellate, lobate frond with at least superficial similarities to the ivesheadiomorph Blackbrookia. The type species is B. alta.

Bulbobaculites felixi[709]

Sp. nov

Valid

Pleş, Bucur & Săsăran

Early Cretaceous (late Aptian)

 Romania

A lituolid foraminifer.

Caelatimurus[710]

Gen. et sp. nov

Valid

Riedman & Porter

Neoproterozoic

Alinya Formation

 Australia

A microfossil. Genus includes new species C. foveolatus.

Chinggiskhaania[711]

Gen. et sp. nov

Dornbos et al.

Ediacaran

Zuun-Arts Formation

 Mongolia

A multicellular benthic alga of uncertain phylogenetic placement. The type species is Chinggiskhaania bifurcata.

Clypeorbis? ultima[712]

Sp. nov

Valid

Schlagintweit, Studeny & Sanders

Late Cretaceous (Maastrichtian)

Kambühel Formation

 Austria

A foraminifer, possibly a member of Clypeorbinae.

Cretaciclavulina[713]

Gen. et sp. nov

Valid

Schlagintweit & Cvetko Tešović

Late Cretaceous (early Campanian)

Gornji Humac Formation
Pučišća Formation

 Croatia

A foraminifer belonging to the superfamily Textulariacea, possibly a member of the family Valvulinidae. The type species is C. gusici.

Cribrionella[714]

Gen. et sp. nov

Valid

Jovanovska et al.

Quaternary

Lake Ohrid

A diatom belonging to the group Thalassiosirales and the family Stephanodiscaceae. The type species is Cribrionella ohridana

Crinisdendrum[715]

Gen. et sp. nov

Valid

Dzik, Baliński & Sun

Ordovician (early Floian)

Fenxiang Formation

 China

An organism of uncertain phylogenetic placement; the authors of its description considered it most likely that it was a relative of pterobranchs. The type species is C. sinicum.

Culcitulisphaera[710]

Gen. et sp. nov

Valid

Riedman & Porter

Neoproterozoic

Alinya Formation

 Australia

A microfossil. Genus includes new species C. revelata.

Delphineis urbinai[716]

Sp. nov

Valid

Gariboldi

Miocene

Pisco Formation

 Peru

A diatom.

Dennisiellinites[717]

Gen. et sp. nov

Valid

Bannister, Conran & Lee

Eocene

Pikopiko Fossil Forest

 New Zealand

A fungus. Genus includes new species D. pikopikoensis .

Diphyes digitum[698]

Sp. nov

Valid

Awad & Oboh-Ikuenobe

Early Paleocene

Gulf of Guinea (Côte D'Ivoire-Ghana Transform Margin)

A dinoflagellate belonging to the group Gonyaulacales.

Discocyclina kutchensis[718]

Sp. nov

Valid

Özcan & Saraswati in Özcan et al.

Eocene (Bartonian)

Fulra Limestone

 India
 Pakistan

A foraminifer belonging to the family Discocyclinidae.

‘Discocyclina’ sulaimanensis[718]

Sp. nov

Valid

Özcan, Ali & Hanif in Özcan et al.

Eocene (Bartonian)

Drazinda Formation

 Pakistan

A foraminifer belonging to the family Discocyclinidae.

Elianella brasiliana[719]

Sp. nov

Valid

Granier & Dias-Brito

Early Cretaceous (Albian)

Riachuelo Formation

 Brazil

A red alga belonging to the group Corallinophycidae, possibly a member of Rhodogorgonales; a species of Elianella.

Ellipsolithus linnertii[707]

Sp. nov

Valid

Lees & Bown

Late Cretaceous (Turonian)

 Tanzania

A microfossil.

Endoceratium immarinum[720]

Sp. nov

Valid

Razumkova

Early Cretaceous (Aptian)

 Russia

A dinoflagellate belonging to the family Ceratiaceae.

Eocladopyxis furculum[698]

Sp. nov

Valid

Awad & Oboh-Ikuenobe

Paleocene to Oligocene

Gulf of Guinea (Côte D'Ivoire-Ghana Transform Margin)

A dinoflagellate belonging to the group Gonyaulacales.

Eohalothece[721]

Gen. et sp. nov

Valid

Strother & Wellman

Precambrian

Cailleach Head Formation
Diabaig Formation
Kinloch Formation
Nonesuch Formation

 United Kingdom
 United States
( Michigan)

A member of Cyanobacteria belonging to the family Chroococcaceae. The type species is Eohalothece lacustrina.

Ercumentina[722]

Gen. et sp. nov

Valid

Serra-Kiel & Vicedo in Serra-Kiel et al.

Paleocene

Jafnayn Formation

 Oman

A foraminifer belonging to the group Miliolida and the superfamily Alveolinoidea. The type species is E. sayqensis.

Gemmaphyton[653]

Gen. et sp. nov

Valid

Wang et al.

Ediacaran

Doushantuo Formation

 China

A macrofossil organism of uncertain phylogenetic placement, possibly an alga. The type species is G. taoyingensis.

Gerontomyces[723]

Gen. et sp. nov

Valid

Poinar

Eocene

Baltic amber

Europe (Baltic Sea coast)

A gilled mushroom. The type species is G. lepidotus.

Glomospirella cantabrica[724]

Sp. nov

Valid

Schlagintweit, Rosales & Najarro

Early Cretaceous (Aptian-Albian)

Las Peñosas Formation
Reocín Formation

 Spain

A foraminifer belonging to the class Tubothalamea, the order Ammodiscida and the family Ammodiscidae.

Gomphosiphon[697]

Gen. et sp. nov

Valid

Liu et al.

Ordovician

Tarim Basin

 China

A possible member of Cyanobacteria. The type species is Gomphosiphon xinjiangensis.

Helicolithus blairiae[725]

Sp. nov

Valid

Kita, Watkins & Bergen

Late Cretaceous (late Santonian)

Niobrara Formation

 United States
( Kansas)

A haptophyte belonging to the family Eiffellithaceae.

Hyalolithus tumescens[726]

Sp. nov

Valid

Abe, Tsutsui & Jordan

Eocene

 Barbados
 United States
( California)

A haptophyte belonging to the group Prymnesiales.

Hypoxylonites foyelensis[727]

Sp. nov

Valid

Bianchinotti, Martínez & Saxena in Martínez et al.

Paleogene (probably Eocene)

Troncoso Formation

 Argentina

A fungus described on the basis of spores.

Idalina grelaudae[704]

Sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Eocene

 Oman
 Yemen

A foraminifer belonging to the group Miliolida and the family Hauerinidae.

Idalina pignattii[704]

Sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Oligocene

Ashawq Formation

 Oman
 Yemen

A foraminifer belonging to the group Miliolida and the family Hauerinidae.

Jimenezberrocosoia[707]

Gen. et comb. et sp. nov

Valid

Lees & Bown

Cretaceous (Aptian-Turonian)

North Sea Basin
 Tanzania

A microfossil. Genus includes "Crucibiscutum" bosunensis Jeremiah (2001), as well as new species J. birchiae.

Jodhpurophycus[728]

Gen. et sp. nov

Valid

Kumar & Ahmad

Ediacaran

Jodhpur Sandstone

 India

A possible alga of uncertain phylogenetic placement. Genus includes new species J. marwarensis.

Kaibabia[729]

Gen. et sp. nov

Valid

Porter & Riedman

Neoproterozoic

Chuar Group

 United States
( Arizona)

A microfossil. Genus includes new species K. gemmulella.

Karenagare[710]

Gen. et sp. nov

Valid

Riedman & Porter

Neoproterozoic

Alinya Formation

 Australia

A microfossil. Genus includes new species K. alinyaensis.

Kokia kayae[707]

Sp. nov

Valid

Lees & Bown

Late Cretaceous (Turonian)

 Tanzania

A microfossil.

Lacazinella rogeri[722]

Sp. nov

Valid

Serra-Kiel & Vicedo in Serra-Kiel et al.

Paleocene

Jafnayn Formation

 Oman

A foraminifer belonging to the group Miliolida and the family Fabulariidae.

Lantianella[730]

Gen. et 2 sp. nov

Valid[731]

Wan et al.

Early Ediacaran

Lantian Formation

 China

An organism of uncertain phylogenetic placement; might be a cnidarian (the possibility considered to be most likely by the authors of its description), or a macroalga. The type species is Lantianella laevis; genus also includes L. annularis.

Latiortenuiphyton[653]

Gen. et sp. nov

Valid

Wang et al.

Ediacaran

Doushantuo Formation

 China

A macroscopic alga of uncertain phylogenetic placement. The type species is L. robusta.

Lindavia cohenii[703]

Sp. nov

Valid

Mohan & Stone in Mohan, Stone & Campisano

Pliocene

Hadar Formation

 Ethiopia

A diatom belonging to the group Thalassiosirales and the family Stephanodiscaceae.

Macetadiscus[704]

Gen. et sp. nov

Valid

Hottinger, Serra-Kiel & Gallardo-Garcia in Serra-Kiel et al.

Eocene

 Oman
 Yemen

A foraminifer belonging to the group Miliolida and the family Soritidae. The type species is M. incolumnatus.

Mallomonas elephantus[732]

Sp. nov

Valid

Siver & Wolfe

Eocene

 Canada

A synurophyte, a species of Mallomonas.

Megacrassispirella[699]

Gen. et comb. nov

Valid

Zhang in Zhang et al.

Permian

Xiala Formation

 China

A foraminifer belonging to the group Miliolata and the family Cornuspiridae. A new genus for "Ammodiscus" xarlashanensis Wang (1986).

Microlepidopalla[729]

Gen. et sp. nov

Valid

Porter & Riedman

Neoproterozoic

Chuar Group

 United States
( Arizona)

A microfossil. Genus includes new species M. mira.

Monodictysporites[733]

Gen. et sp. nov

Valid

Klymiuk

Eocene (YpresianLutetian)

Allenby Formation

 Canada
( British Columbia)

A fungus described on the basis of spores. Genus includes new species M. princetonensis.

Morgensternia[710]

Gen. et sp. nov

Valid

Riedman & Porter

Neoproterozoic

Alinya Formation

 Australia

A microfossil. Genus includes new species M. officerensis.

Nannoconus funiculus[707]

Sp. nov

Valid

Lees & Bown

Late Cretaceous (Turonian)

 Tanzania

A microfossil.

Nodocantabricus[734]

Gen. et sp. nov

Valid

Rigaud & Schlagintweit

Late Cretaceous (Cenomanian)

Altamira Formation
Bielba Formation

 Spain

A foraminifer belonging to the family Polymorphinidae. Genus includes new species N. duplexmurus.

Nyktericysta sibirica[720]

Sp. nov

Valid

Razumkova

Early Cretaceous (Aptian)

 Russia

A dinoflagellate belonging to the family Ceratiaceae.

Omanodiscus[704]

Gen. et sp. nov

Valid

Hottinger, Serra-Kiel & Gallardo-Garcia in Serra-Kiel et al.

Eocene

 Oman

A foraminifer belonging to the group Miliolida and the family Soritidae. The type species is O. tenuissimus.

Ortonellopsis[701]

Gen. et sp. et comb. nov

Valid

Vachard & Cózar in Vachard et al.

Silurian-Triassic

 Canada
 France
 Greece
 Spain

A member of Cyanobacteria belonging to the family Garwoodiaceae. The type species is O. laxa; genus also includes "Ortonella" mansellensis (Poncet, 1986) and "Ortonella" myrae Rácz (1964).

Paleocadus[735]

Gen. et sp. nov

Valid

Poinar

Cretaceous

Burmese amber

 Myanmar

A member of Eccrinales belonging to the family Eccrinaceae. The type species is P. burmiticus.

Paragraptobranca[653]

Gen. et sp. nov

Valid

Wang et al.

Ediacaran

Doushantuo Formation

 China

A macrofossil organism of uncertain phylogenetic placement, might be a macroalga or an animal. The type species is P. curvus.

Perissothallus dekovensis[736]

Sp. nov

Valid

Pšenička & Krings

Carboniferous (Gzhelian)

Slaný Formation

 Czech Republic

An alga of uncertain phylogenetic placement, a species of Perissothallus.

Petrarhabdus? kirenii[707]

Sp. nov

Valid

Lees & Bown

Late Cretaceous (Turonian)

 Tanzania

A microfossil.

Piyuania[730]

Gen. et sp. nov

Valid[731]

Wan et al.

Early Ediacaran

Lantian Formation

 China

An organism of uncertain phylogenetic placement, possibly a cnidarian-grade animal. The type species is P. cyathiformis.

Pluricellaesporites patagonicus[727]

Sp. nov

Valid

Bianchinotti, Martínez & Saxena in Martínez et al.

Paleogene

 Argentina
 Turkey

A fungus described on the basis of spores resembling those of members of the genus Bactrodesmium Cooke.

Plumeropriscum[708]

Gen. et sp. nov

Valid

Mason & Narbonne

Ediacaran

 Canada
( Newfoundland and Labrador)

A mop-like rangeomorph. The type species is P. hofmanni.

Proaulopora pachydermatica[697]

Sp. nov

Valid

Liu et al.

Ordovician

Tarim Basin

 China

A possible member of Cyanobacteria, a species of Proaulopora.

Pseudoaccordiella[704]

Gen. et sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Eocene

 Oman
 Yemen

A foraminifer belonging to the group Globothalamea and the family Pfenderinidae. The type species is P. ayaki.

Pseudodoushantuophyton[653]

Gen. et sp. nov

Valid

Wang et al.

Ediacaran

Doushantuo Formation

 China

A probably a macroalga. The type species is P. wenghuiensis.

Pseudolituonella robineti[704]

Sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Eocene

Aydim Formation

 Oman
 Yemen

A foraminifer belonging to the group Globothalamea and the family Coskinolinidae.

Pseudonumoloculina kalantarii[696]

Sp. nov

Valid

Schlagintweit & Rashidi

Late Cretaceous (Maastrichtian)

Tarbur Formation

 Iran

A foraminifer belonging to the family Hauerinidae.

Pseudorhapydionina bilottei[737]

Sp. nov

Valid

Consorti, Boix & Caus

Late Cretaceous (Santonian)

La Cova Unit

 Spain

A foraminifer belonging to the group Miliolida and the family Praerhapydioninidae.

Qianchuania[730]

Gen. et sp. nov

Valid[731]

Wan et al.

Early Ediacaran

Lantian Formation

 China

An organism of uncertain phylogenetic placement, possibly a cnidarian-grade animal. The type species is Q. fusiformis.

Rogerella[704]

Gen. et sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Eocene

Aydim Formation

 Oman
 Yemen

A foraminifer belonging to the group Globothalamea and the family Orbitolinidae. The type species is R. aydimi.

Rossanella[738]

Gen. et sp. nov

Valid

Rigaud & Blau

Early Jurassic (HettangianSinemurian)

Schnöll Formation

 Austria

A foraminifer belonging to the group Robertinida and the family Conorboididae. The type species is R. martinii.

Rothpletzella longita[697]

Sp. nov

Valid

Liu et al.

Ordovician

Tarim Basin

 China

A microorganism of uncertain phylogenetic placement, a species of Rothpletzella.

Socotraella[704]

Gen. et sp. nov

Valid

Gallardo-Garcia & Serra-Kiel in Serra-Kiel et al.

Oligocene

Ashawq Formation
Aydim Formation

 Oman
 Yemen

A foraminifer belonging to the group Globothalamea and the family Valvulinidae. The type species is S. ashawqi.

Sphenolithus pospichalii[739]

Sp. nov

Valid

Jiang et al.

Early Miocene

Northern shelf of the South China Sea

A microfossil.

Spirolina? farsiana[696]

Sp. nov

Valid

Schlagintweit & Rashidi

Late Cretaceous (Maastrichtian)

Tarbur Formation

 Iran

A foraminifer belonging to the group Miliolida and the superfamily Soritoidea.

Surninia[740]

Gen. et sp. nov

Valid

Kolosov

Ediacaran

 Russia

A microorganism. Genus includes new species S. implicata.

Tectatodinium nigeriaense[698]

Sp. nov

Valid

Awad & Oboh-Ikuenobe

Paleocene

Gulf of Guinea (Côte D'Ivoire-Ghana Transform Margin)
 Nigeria

A dinoflagellate belonging to the group Gonyaulacales.

Tongrenphyton[653]

Gen. et sp. nov

Valid

Wang et al.

Ediacaran

Doushantuo Formation

 China

A probably an eukaryotic alga. The type species is T. komma.

Trewinomyces[741]

Gen. et sp. nov

Valid

Krings, Taylor & Martin

Early Devonian

Rhynie chert

 United Kingdom

A fungus of uncertain phylogenetic placement, showing similarities to members of the genera Macrochytrium (Chytridiomycota) and Blastocladiella (Blastocladiomycota). The type species is Trewinomyces annulifer.

Trochoguembelitria liuae[742]

Sp. nov

Valid

Arenillas, Arz & Náñez

Paleocene (early Danian)

El Haria Formation

 Tunisia

A foraminifer belonging to the group Heterohelicacea and the family Guembelitriidae.

Trochoguembelitria olssoni[742]

Sp. nov

Valid

Arenillas, Arz & Náñez

Paleocene (early Danian)

El Haria Formation

 Tunisia

A foraminifer belonging to the group Heterohelicacea and the family Guembelitriidae.

Velleditsiella[738]

Gen. et 2 sp. nov

Valid

Rigaud & Blau

Early Jurassic (HettangianSinemurian)

Schnöll Formation

 Austria

A foraminifer belonging to the group Robertinida and the family Trochosiphoniidae. The type species is V. felicitaszae; genus also includes V. spinaferra.

Vendophycus[728]

Gen. et 2 sp. nov

Valid

Kumar & Ahmad

Ediacaran

Jodhpur Sandstone

 India

A possible alga of uncertain phylogenetic placement. Genus includes new species V. rajasthanensis and V. sursagarensis.

Vidalopalla[710]

Gen. et comb. nov

Valid

Riedman & Porter

Neoproterozoic

 Norway

A microfossil. Genus includes "Kildinosphaera" verrucata Vidal in Vidal & Siedlecka (1983).

Volleyballia[729]

Gen. et sp. nov

Valid

Porter & Riedman

Neoproterozoic

Chuar Group

 United States
( Arizona)

A microfossil. Genus includes new species V. dehlerae.

Xianella[743]

Gen. et sp. nov

Valid

Lee & Riding

Middle–Late Ordovician

 China

A member of Cyanobacteria. The type species is X. hongii.

Xiuningella[730]

Gen. et sp. nov

Valid[731]

Wan et al.

Early Ediacaran

Lantian Formation

 China

An organism of uncertain phylogenetic placement; might be a worm-like animal or an alga. The type species is X. rara.

Zuunartsphyton[711]

Gen. et sp. nov

Dornbos et al.

Ediacaran

Zuun-Arts Formation

 Mongolia

A multicellular benthic alga of uncertain phylogenetic placement. The type species is Zuunartsphyton delicatum.

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