2013 in paleontology

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Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.^[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2013.

Plants

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Atli^[2]	Gen. et sp. nov.	Valid	Smith et al.	Late Cretaceous (Campanian)		Canada	A ranunculalean known from a stem with lianoid anatomy. The type specimen is Atli morinii.
Bambusiculmus^[3]	Gen. et 2 sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo. The type specimen is Bambusiculmus angustus; genus also includes Bambusiculmus latus.
Bambusium angustifolia^[3]	Sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo, a species of Bambusium.
Bambusium latifolia^[3]	Sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo, a species of Bambusium.
Bombacacioxylon^[4]	Gen. et sp. nov.	Valid	Mehrotra et al.	Late Miocene to early Pliocene	Tipam Group	India	A possible relative of Bombax. The type specimen is Bombacacioxylon tertiarum.
Firmiana oligocenica^[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Firmiana.
Guadua morronei^[6]	Sp. nov.	Valid	Brea, Zucol & Franco	Pliocene–Pleistocene	Ituzaingó Formation	Argentina	A bamboo, a species of Guadua.
Iterophyllum^[7]	Gen. et sp. nov.	Valid	Barral et al.	Early Cretaceous (late Barremian)		Spain	A basal eudicot. The type specimen is Iterophyllum lobatum.
Jaguariba wiersemana^[8]	Gen. nov. et sp. nov.	Valid	Coiffard, Mohr & Bernardes-de-Oliveira	Early Cretaceous	Crato Formation	Brazil	Fossil Nymphaeales.
Laurelia otagoensis^[9]	Sp. nov	Valid	Conran, Bannister & Lee	Early Miocene		New Zealand	A species of Laurelia.
Palmoxylon vastanensis^[10]	Sp. nov.	Valid	Prasad et al.	Eocene	Cambay Shale Formation	India	A palm tree, a species of Palmoxylon.
Paranephelium makumensis^[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Paranephelium.
Poeciloneuron preindicum^[11]	Sp. nov.	Valid	Srivastava & Mehrotra	Late Oligocene		India	A species of Poeciloneuron.
Potomacapnos^[12]	Gen. et sp. nov.	Valid	Jud & Hickey	Early Cretaceous (Aptian)	Potomac Group	United States	A close relative of ranunculalean eudicots. The type specimen is Potomacapnos apeleutheron.
Pterygota palaeoalata^[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Pterygota.
Sapindus palaeoemarginatus^[5]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Sapindus.
Spixiarum kipea^[13]	Gen. nov. et sp. nov.	Valid	Coiffard, Mohr & Bernardes-de-Oliveira	Early Cretaceous	Crato Formation	Brazil	Fossil Aroid.
Virola dominicana^[14]	Sp. nov.	Valid	Poinar & Steeves	Cenozoic (Eocene or Miocene)		Dominican Republic	A member of Myristicaceae found in Dominican amber, a species of Virola.

Other seed plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Bancroftiastrobus^[15]	Gen. et sp. nov.	Valid	Rothwell et al.	Late Jurassic (early Kimmeridgian)	Kimmeridge Clay Formation	United Kingdom	A member of Coniferales of uncertain phylogenetic placement. The type specimen is Bancroftiastrobus digitata.
Chengia^[16]	Gen. et sp. nov.	Valid	Yang, Lin & Wang	Early Cretaceous (Aptian)	Yixian Formation	China	A member of Ephedraceae. The type specimen is Chengia laxispicata.
Cunninghamia taylorii^[17]	Sp. nov.	Valid	Serbet, Bomfleur & Rothwell	Late Cretaceous (Campanian)	Horseshoe Canyon Formation	Canada	A species of Cunninghamia.
Donponoxylon^[18]	Gen. et 2 sp. nov.	Valid	Tidwell, Britt & Wright	Middle to Late Jurassic		Australia New Zealand	A seed plant of uncertain phylogenetic placement. The type specimen is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii.
Dordrechtites arcanus^[19]	Sp. nov.	Valid	Bergene, Taylor & Taylor	Middle Triassic (Anisian)	Fremouw Formation	Antarctica	A gymnosperm of uncertain phylogenetic placement.
Friedsellowia gracilifolia^[20]	Gen. nov. et sp. nov.	Valid	Löwe et al.	Early Cretaceous	Crato Formation	Brazil	A gnetophyte. The type of a new genus.
Itajuba^[21]	Gen. et sp. nov	Valid	Ricardi-Branco et al.	Early Cretaceous (Aptian or Albian)	Crato Member of the Santana Formation	Brazil	A member of Gnetales. The type specimen is Itajuba yansanae.
Pagiophyllum heshanense^[22]	Sp. nov.	Valid	Liu & Yao	Permian		China	A species of Pagiophyllum.
Pararaucaria delfueyoi^[23]	Sp. nov.	Valid	Escapa et al.	Late Jurassic	Cañadón Calcáreo Formation	Argentina	A member of Coniferales belonging to the family Cheirolepidiaceae, a species of Pararaucaria.
Prumnopitys anglica^[24]	Sp. nov.	Valid	Greenwood, Hill & Conran	Eocene	Branksome Formation	United Kingdom	A species of Prumnopitys.
Schizolepidopsis canicularis^[25]	Sp. nov.	Valid	Leslie et al.	Early Cretaceous (probably Aptian-Albian)	Tevshiin Govi Formation	Mongolia	A member of Coniferales, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis.
Stephanospermum braidwoodensis^[26]	Sp. nov.	Valid	Spencer, Hilton & Sutton	Carboniferous	Mazon Creek fossil beds	United States	A member of Medullosales, a species of Stephanospermum.
Stephanospermum shuichengensis^[27]	Sp. nov.	Valid	Spencer et al.	Permian (Lopingian)	Wangjiazhai Formation	China	A member of Medullosales, a species of Stephanospermum.

Other plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Ashicaulis beipiaoensis^[28]	Sp. nov	Valid	Tian et al.	Middle Jurassic	Tiaojishan Formation	China	A member of Osmundaceae, a species of Ashicaulis.
Bryiidites^[29]	Gen. et sp. nov.	Valid	Barclay et al.	Late Cretaceous (late Cenomanian)	Middle Dakota Formation	United States	A member of Bryidae of uncertain phylogenetic placement. The type specimen is Bryiidites utahensis.
Franhueberia^[30]	Gen. et sp. nov.	Valid	Hoffman & Tomescu	Devonian (Emsian)	Battery Point Formation	Canada	A basal member of Euphyllophytina. The type specimen is Franhueberia gerriennei.
Palaeosorum^[31]	Gen. et sp. nov	Valid	Jacques & Zhou in Jacques, Su & Zhou	Miocene	Dajie Formation	China	A member of Polypodiaceae. The type specimen is Palaeosorum ellipticum.
Yavanna^[32]	Gen. et sp. nov	Valid	Vera	Early Cretaceous (Aptian)	Cerro Negro Formation	Antarctica (Livingston Island)	A member of Cyatheales of uncertain phylogenetic placement. The type specimen is Yavanna chimaerica.

Arthropods

Main article: 2013 in arthropod paleontology

Bryozoans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Adlatipora^[33]	Gen. et sp. nov	Valid	Gautier, Jackson & McKinney	Permian		United States	An acanthocladiid bryozoan. The type species is Adlatipora fossulata.
Basyaylella^[34]	Gen. et sp. nov	Valid	Zágoršek & Gordon	Late Tortonian	Mut Basin	Turkey	An ascophoran bryozoan. Its type species is Basyaylella elsae.
Charixa goshouraensis^[35]	Sp. nov	Valid^[36]	Dick et al.	Cretaceous (late Albian or early Cenomanian)		Japan	A malacostegan, a species of Charixa.
Conopeum wilsoni^[37]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Conopeum.
Haplostoechios^[35]	Gen. et 2 sp. nov	Valid^[36]	Dick et al.	Cretaceous (late Albian or early Cenomanian)		Japan	A malacostegan. Genus contains two species: H. hayamiae and H. clusum.
Heloclema uralicum^[38]	Sp. nov	Valid	Tolokonnikova	Early Carboniferous		Russia	A species of Heloclema.
Monoceratopora whybrowi^[37]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Monoceratopora.
Ostrovskia^[34]	Gen. et sp. nov	Valid	Zágoršek & Gordon	Late Tortonian	Mut Basin	Turkey	An ascophoran bryozoan. Its type species is Ostrovskia triforamina.
Primorella kodinkensis^[38]	Sp. nov	Valid	Tolokonnikova	Early Carboniferous		Russia	A species of Primorella.
Tyloporella smithi^[37]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Tyloporella.
Wilbertopora cheethami^[37]	Sp. nov	Valid	Di Martino & Taylor	Late Cretaceous (Late Campanian or Maastrichtian)	Simsima Formation	United Arab Emirates–Oman border region	A species of Wilbertopora.

Brachiopods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Arenorthis paranaensis^[39]	Sp. nov	Valid	Benedetto, Halpern & Inchausti	Late Ordovician (Hirnantian)	Eusebio Ayala Formation	Paraguay	A species of Arenorthis.
Askepasma saproconcha^[40]	sp nov	Valid	Topper et al.	Lower Cambrian	Wirrapowie Limestone	Australia	A member of Paterinata, a species of Askepasma.
Biseptata^[41]	Gen. et sp. nov	Valid	Madison	Ordovician		Russia	A member of Strophomenida. The type species is Biseptata briani.
Menathyris^[42]	Gen. et sp. nov	Valid	Feldman	Middle Triassic (early Ladinian)	Saharonim Formation	Israel	A zeilleriid terebratulid. The type species is Menathyris wilsoni.
Neochonetes (Huangichonetes) archboldi^[43]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Neochonetes semicircularis^[43]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Neochonetes (Sommeriella) rectangularis^[43]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Neochonetes (Sommeriella) waterhousei^[43]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Neochonetes.
Plectothyrella? itacurubiensis^[39]	Sp. nov	Valid	Benedetto, Halpern & Inchausti	Late Ordovician (Hirnantian)	Eusebio Ayala Formation	Paraguay	Possibly a species of Plectothyrella.
Proclinorthis^[44]	Gen. et sp. nov	Valid	Colmenar, Villas & Vizcaïno	Late Ordovician	Glauzy Formation	France	A platyorthid. The type species is Proclinorthis vailhanensis.
Pseudocrania insperata^[45]	Sp. nov	Valid	Bassett et al.	Middle Ordovician (Darriwilian)	Lashkarak Formation	Iran	A member of Craniida, a species of Pseudocrania.
Sphenope^[46]	Gen. et comb. et sp. nov	Valid	Vörös	Late Jurassic to Early Cretaceous (Tithonian to Berriasian)		Hungary Italy	A pygopid terebratulid. A new genus for "Terebratula" misilmerensis Gemmellaro (1871); genus also contains the new species Sphenope bifida.
Tethyochonetes cheni^[43]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Tethyochonetes.
Tethyochonetes sheni^[43]	Sp. nov	Valid	Zhang et al.	Late Permian (Changhsingian)		China	A member of Rugosochonetidae, a species of Tethyochonetes.

Molluscs

Main article: 2013 in molluscan paleontology

Echinoderms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Abertella miskellyi^[47]	Sp. nov	Valid	Kroh et al.	Miocene	Camarones Formation	Argentina	An abertellid sand dollar, a species of Abertella.
Alternacantha^[48]	Gen. et sp. nov	Valid	Thuy & Meyer	Middle Jurassic		Switzerland	An ophiacanthid brittle star. The type species is Alternacantha occulta.
Aragocystites^[49]	Gen. et sp. nov	Valid	Zamora	Middle Cambrian	Murero Formation	Spain	An edrioasteroid. The type species is Aragocystites belli.
Bathyovulaster^[50]	Gen. et sp. nov	Valid	Smith	Santonian		Italy	A spatangoid. The type species is Bathyovulaster disjunctus.
Bellastrella^[51]	Gen. et sp. nov	Valid	Müller, Hahn and Bohatý	Middle Devonian		Germany	An agelacrinitid edrioasteroid. The type species is Bellastrella eifeliana.
Ctenocrinus branisai^[52]	Sp. nov	Valid	Thompson, Ausich & Smith	Early Devonian (Emsian)	Belèn Formation	Bolivia	A crinoid, a species of Ctenocrinus.
Cyclolampas altus^[53]	Sp. nov	Valid	Saucede et al.	Middle Jurassic (late Callovian)		France	An echinoid, a species of Cyclolampas.
Eozonella^[54]	Gen. et sp. nov	Valid	Thuy, Marty & Comment	Late Jurassic (late Oxfordian)		Switzerland	An ophiolepidid brittle star. The type species is Eozonella bergeri.
Graciacystis^[55]	Gen. et sp. nov	Valid	Zamora, Rahman & Smith	Middle Cambrian		Spain	A basal cinctan echinoderm. The type species is Graciacystis ambigua.
Griphocrinus pirovanoi^[52]	Sp. nov	Valid	Thompson, Ausich & Smith	Early Devonian (Emsian)	Icla Formation	Bolivia	A crinoid, a species of Griphocrinus.
Hanshessia^[48]	Gen. et sp. nov	Valid	Thuy & Meyer	Middle Jurassic		Switzerland	An ophiacanthid brittle star. The type species is Hanshessia trochitophila.
Juracantha^[54]	Gen. et sp. nov	Valid	Thuy, Marty & Comment	Late Jurassic (late Oxfordian)		Switzerland	An ophiacanthid brittle star. The type species is Juracantha hottingeri.
Kosachenkoastrus^[56]	Gen. et sp. nov	Valid	Rozhnov	Middle Ordovician		Russia	A member of Parablastoidea. The type species is Kosachenkoastrus volkhovensis.
Lutocrinus^[52]	Gen. et sp. nov	Valid	Thompson, Ausich & Smith	Early Devonian (Emsian)	Icla Formation	Bolivia	A crinoid. The type species is Lutocrinus boliviaensis.
Megaradialocrinus dorotheakochae^[57]	Sp. nov	Valid	Bohatý & Hein	Devonian		Germany	A camerate crinoid belonging to the group Monobathrida and the family Hexacrinitidae.
Ophiotholia aurora^[48]	Sp. nov	Valid	Thuy & Meyer	Middle Jurassic		Switzerland	An ophiomycetid brittle star, a species of Ophiotholia.
Regulaecystis testudineus^[58]	Sp nov	Valid	Nardin & Bohatý	Devonian (Eifelian)		Germany	A blastozoan belonging to the family Pleurocystitidae.
Rhenopyrgus piojoensis^[59]	Sp nov	Valid	Sumrall, Heredia, Rodríguez & Mestre	Silurian (lower Ludlow)	Los Espejos Formation	Argentina	A rhenopyrgid edrioasteroid, a species of Rhenopyrgus.
Simakocrinus^[60]	Gen. et 2 sp. nov.	Valid	Prokop	Devonian (Pragian, Emsian and Eifelian)		Czech Republic	A crinoid belonging to the group Disparida, possibly a member of the family Myelodactylidae. The type species is Simakocrinus facilis sp. nov (col.); genus also contains Simakocrinus diligens sp. nov (col.).

Conodonts

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acanthodus humachensis^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acanthodus.
Acanthodus raqueli^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acanthodus.
Acodus primitivus^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acodus.
Acuminatella^[62]	Gen. et 2 sp. nov	Valid	Orchard	Late Triassic (late Carnian to early Norian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. The type species is Acuminatella acuminata; genus also contains Acuminatella angusta.
Bispathodus ultimus bartzschi^[63]	Subsp nov	Valid	Kononova & Weyer	Devonian (late Famennian)		Germany	A subspecies of Bispathodus ultimus.
Carnepigondolella eozoae^[62]	Sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Ludington Formation	Canada	A gondolellid ozarkodinid, a species of Carnepigondolella.
Carnepigondolella medioconstricta^[62]	Sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Ludington Formation	Canada	A gondolellid ozarkodinid, a species of Carnepigondolella.
Furnishina leei^[64]	Sp. nov	Valid	Lee	Cambrian	Machari Formation	South Korea
Histiodella wuhaiensis^[65]	Sp. nov	Valid	Wang et al.	Ordovician	Lower Klimoli Formation	China
Idiognathodus craticulinodosus^[66]	Sp. nov	Valid	Ishida, Suzuki & Inada	Carboniferous (middle–late Bashkirian to early Moscovian)	Ko-yama Limestone Group	Japan
Kallidontus gondwanicus^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Kallidontus.
Kraussodontus^[62]	Gen. et sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. The type species is Kraussodontus peteri.
Mesogondolella daheshenensis^[67]	Sp nov	Valid	Wang in Zhou et al.	Permian (probably Wordian)	Daheshen Formation	China	A species of Mesogondolella.
Mesogondolella subgracilis^[67]	Sp nov	Valid	Wang in Zhou et al.	Permian (probably Wordian)	Daheshen Formation	China	A species of Mesogondolella.
Muellerilepis^[68]	Nom. nov	Valid	Bardashev & Bardasheva	Middle Devonian (Givetian)		Tajikistan	A replacement name for Muellerina Bardashev & Bardasheva (2012) (preoccupied).
Naimanodus^[69]	Gen. et sp. nov	Valid	Tolmacheva	Middle Ordovician		Kazakhstan Kyrgyzstan Russia	The type species is Naimanodus degtyarevi.
Neopolygnathus margaretae^[63]	Sp nov	Valid	Kononova & Weyer	Devonian (late Famennian)		Germany	A species of Neopolygnathus.
Notiodella^[70]	Gen. et sp. nov	Valid	Aldridge et al.	Late Ordovician	Soom Shale Member, Cedarberg Formation	South Africa	A member of Prioniodontida belonging to the family Balognathidae. The type species is Notiodella keblon.
Palmatolepis khaensis^[71]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Palmatolepis marki^[71]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Palmatolepis subperlobata lapoensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Parapetella^[62]	Gen. et sp. nov	Valid	Orchard	Late Triassic (late Carnian to early Norian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. The type species is Parapetella prominens.
Polygnathus burretti^[71]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus communis hanensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus communis namdipensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus communis phaphaensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus crassulus salapensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus extralobatus phoensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Polygnathus hojedki^[72]	Sp. nov	Valid	Gholamalian et al.	Devonian		Iran	A species of Polygnathus.
Polygnathus pseudocostatus^[73]	Sp nov	Valid	Klapper & Vodrážková	Devonian		United States	A species of Polygnathus.
Primatella^[62]	Gen. et comb. et 2 sp. nov	Valid	Orchard	Late Triassic (late Carnian to early Norian)	Pardonet Formation	Canada	A gondolellid ozarkodinid. A new genus for "Epigondolella" primitia Mosher (1970); genus also contains new species Primatella asymmetrica and Primatella conservativa.
Pseudopolygnathus granulosus laepensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Pseudopolygnathus granulosus salawinensis^[71]	Subsp. nov	Valid	Savage	Late Devonian		Thailand
Pseudopolygnathus maepoensis^[71]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Quadralella^[62]	Gen. et sp. nov	Valid	Orchard	Late Triassic (late Carnian)	Ludington Formation	Canada	A gondolellid ozarkodinid. The type species is Quadralella lobata.
Siphonodella banraiensis^[71]	Sp. nov	Valid	Savage	Late Devonian		Thailand
Tilcarodus^[61]	Gen. et comb. nov	Valid	Zeballo & Albanesi	Early Ordovician	Rupasca Formation	Argentina	A new genus for "Utahconus" humahuacensis Albanesi and Aceñolaza (2005).
Trapezognathus? primitivus^[74]	Sp. nov	Valid	Voldman, Albanesi & Zeballo in Voldman et al.	Early Ordovician	Acoite Formation	Argentina	Originally described as a possible species of Trapezognathus; subsequently made the type species of a separate genus Zentagnathus.^[75]
Utahconus purmamarcensis^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Utahconus scandodiformis^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Utahconus tortibasis^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Variabiloconus crassus^[61]	Sp nov	Valid	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Variabiloconus.

Fishes

Main article: 2013 in fish paleontology

Amphibians

Newly named temnospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Broiliellus reiszi^[76]	Sp. nov	Valid	Holmes, Berman & Anderson	Early Permian	El Cobre Canyon Formation	United States	A dissorophid, a species of Broiliellus.
Parotosuchus ptaszynskii^[77]	Sp nov	Valid	Sulej & Niedźwiedzki	Late Olenekian		Poland	A species of Parotosuchus.
Reiszerpeton^[78]	Gen. et sp. nov	Valid	Maddin et al.	Early Permian	Archer City Formation	United States	A dissorophid. The type species is Reiszerpeton renascentis.
Scapanops^[79]	Gen. et sp. nov	Valid	Schoch & Sues	Early Permian	Nocona Formation	United States	A dissorophid. The type species is Scapanops neglecta.
Tersomius dolesensis^[80]	Sp nov	Valid	Anderson & Bolt	Early Permian		United States	A species of Tersomius.
Trimerorhachis greggi^[81]	Sp nov	Valid	Milner & Schoch	Early Permian		United States	A species of Trimerorhachis.

Newly named lepospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Batropetes niederkirchensis^[82]	Sp. nov	Valid	Glienke	Early Permian (Rotliegend)	Saar–Nahe Basin	Germany	A species of Batropetes.
Huskerpeton^[83]	Gen. et sp. nov	Valid	Huttenlocker et al.	Early Permian	Eskridge Formation	United States	A recumbirostran. The type species is Huskerpeton englehorni.
Proxilodon^[83]	Gen. et comb. nov	Valid	Huttenlocker et al.	Early Permian	Speiser Formation	United States	A recumbirostran, a new genus for "Euryodus" bonneri.

Newly named lissamphibians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Aerugoamnis^[84]	Gen. et sp. nov	Valid	Henrici, Báez & Grande	Early Eocene (Wasatchian)	Green River Formation	United States	A relative of parsley frogs. The type species is Aerugoamnis paulus.
Eobarbourula^[85]	Gen. et sp. nov	Valid	Folie et al.	Early Eocene	Cambay Shale Formation	India	A bombinatorid. Its type species is Eobarbourula delfinoi.
Gracilibatrachus^[86]	Gen. et sp. nov	Valid	Báez	Early Cretaceous (late Barremian)		Spain	A relative of the family Pipidae. The type species is Gracilibatrachus avallei.
Iberobatrachus^[86]	Gen. et sp. nov	Valid	Báez	Early Cretaceous (late Barremian)		Spain	A close relative of the genus Discoglossus. The type species is Iberobatrachus angelae.
Ichthyosaura randeckensis^[87]	Sp. nov	Valid	Schoch & Rasser	Miocene		Germany	A newt related to the Alpine newt.
Indorana^[85]	Gen. et sp. nov	Valid	Folie et al.	Early Eocene	Cambay Shale Formation	India	A possible rhacophorid. Its type species is Indorana prasadi.
Leiopelma acricarina^[88]	Sp. nov	Valid	Worthy et al.	Early Miocene		New Zealand	A member of Leiopelmatidae.
Leiopelma miocaenale^[88]	Sp. nov	Valid	Worthy et al.	Early Miocene		New Zealand	A member of Leiopelmatidae.
Liaobatrachus zhaoi^[89]	Sp. nov	Valid	Dong et al.	Early Cretaceous (Barremian)	Yixian Formation	China	An anuran of uncertain phylogenetic placement, a species of Liaobatrachus.
Paranecturus^[90]	Gen. et sp. nov	Valid	Demar	Late Cretaceous (Maastrichtian)	Hell Creek Formation	United States	A mudpuppy. The type species is Paranecturus garbanii.
Ukrainurus^[91]	Gen. et sp. nov	Valid	Vasilyan et al.	Miocene		Ukraine	A relative of cryptobranchids. The type species is Ukrainurus hypsognathus.
Wesserpeton^[92]	Gen. et sp. nov	Valid	Sweetman & Gardner	Barremian	Wessex Formation	England	An albanerpetontid.

Turtles

Newly named turtles

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Adocus inexpectatus^[93]	Sp. nov	Valid	Danilov et al.	Late Eocene	Youganwo Formation	China	An adocid, a species of Adocus.
Atoposemys^[94]	Gen. et sp. nov	Valid	Hutchison	Early Paleocene (Puercan)	Fort Union Formation	United States	A trionychid. The type species is Atoposemys entopteros.
Bairdemys healeyorum^[95]	Sp. nov	Valid	Weems & Knight	Late Oligocene	Chandler Bridge Formation	United States	A podocnemidid pleurodiran, a species of Bairdemys.
Basilemys gaffneyi^[96]	Sp. nov	Valid	Sullivan, Jasinski & Lucas	Late Campanian		United States	A nanhsiungchelyid turtle, a species of Basilemys.
Brachyopsemys^[97]	Gen. et sp. nov	Valid	Tong & Meylan	Early Paleocene	Ouled Abdoun Basin	Morocco	A cryptodiran closely related to Sandownia. The type species is Brachyopsemys tingitana.
Camerochelys^[98]	Gen. et sp. nov	Valid	Pérez-García & Murelaga	Early Cretaceous (Hauterivian or Barremian)	Cameros Basin	Spain	A member of Pan-Cryptodira (the clade containing living cryptodirans and all turtles sharing a more recent common ancestor with them than with pleurodirans), possibly a xinjiangchelyid. The type species is Camerochelys vilanovai.
Cardichelyon^[94]	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	A platysternid. The type species is Cardichelyon rogerwoodi.
?Centrochelys marocana^[99]	Sp. nov	Valid	Gmira et al.	Pliocene		Morocco	A tortoise, possibly a species of Centrochelys.
Changmachelys^[100]	Gen. et sp. nov	Valid	Brinkman et al.	Early Cretaceous	Xiagou Formation	China	A member of (likely paraphyletic) group "Macrobaenidae". The type species is Changmachelys bohlini.
Chelodina (Chelodina) murrayi^[101]	Sp. nov	Valid	Yates	Late Miocene	Waite Formation	Australia	A chelid, a species of Chelodina.
Chelonoidis lutzae^[102]	Sp. nov	Valid	Zacarías et al.	Late Pleistocene (58–22 ka)		Argentina	A tortoise, a species of Chelonoidis.
Chrysemys isoni^[103]	Sp. nov	Valid	Weems & George	Miocene	Calvert Formation	United States	A relative of the painted turtle.
Cuora chiangmuanensis^[104]	Sp. nov	Valid	Naksri et al.	Late Middle or early Late Miocene		Thailand	An Asian box turtle.
Floridemys hurdi^[103]	Sp. nov	Valid	Weems & George	Miocene	Calvert Formation	United States	A tortoise, a species of Floridemys.
Kappachelys^[105]	Gen. et sp. nov	Valid	Hirayama, Isaji & Hibino	Early Cretaceous	Akaiwa Formation	Japan	A relative of trionychids. The type species is Kappachelys okurai.
Laganemys^[106]	Gen. et sp. nov	Valid	Sereno & ElShafie	Aptian or Albian	Elrhaz Formation	Niger	An araripemydid pleurodiran. The type species is Laganemys tenerensis.
Neochelys liriae^[107]	Sp. nov	Valid	Pérez-García & de Lapparent de Broin	Eocene (Ypresian)		France	A podocnemidid, a species of Neochelys.
Neurankylus lithographicus^[108]	Sp. nov	Valid	Larson et al.	Santonian	Milk River Formation	Canada	A neurankyline baenid, a species of Neurankylus.
Nostimochelone^[109]	Gen. et sp. nov	Valid	Georgalis et al.	Early Miocene	Zeugostasion Formation	Greece	A pleurodiran, a member of Podocnemidoidea. The type species is Nostimochelone lampra.
Ocadia tanegashimensis^[110]	Sp. nov	Valid	Takahashi et al.	Early middle Miocene	Kawachi Formation	Japan	A geoemydid, a species of Ocadia.
Ocepechelon^[111]	Gen. et sp. nov	Valid	Bardet et al.	Late Cretaceous (late Maastrichtian)	Oulad Abdoun Basin	Morocco	A relative of dermochelyids and protostegids. The type species is Ocepechelon bouyai.
Paramongolemys^[112]	Gen. et sp. nov	Valid	Danilov & Sukhanov	Late Paleocene	Naranbulak Formation	Mongolia	A basal member of Testudinoidea. The type species is Paramongolemys khosatzkyi.
Planetochelys dithyros^[94]	Sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	A relative of trionychids, a species of Planetochelys.
Psilosemys^[94]	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	The earliest North American emydid. The type species is Psilosemys wyomingensis.
Scabremys^[96]	Gen. et comb. nov	Valid	Sullivan, Jasinski & Lucas	Late Campanian		United States	A baenid cryptodiran, a new genus for "Baena" ornata Gilmore (1935).
Shandongemys^[113]	Gen. et sp. nov	Valid	Li et al.	Late Cretaceous		China	A lindholmemydid testudinoid. The type species is Shandongemys dongwuica.
Sinemys brevispinus^[114]	Sp. nov	Valid	Tong & Brinkman	Early Cretaceous	Luohandong Formation	China	A sinemydid cryptodiran, a species of Sinemys.
Spoochelys^[115]	Gen. et sp. nov	Valid	Smith & Kear	Early or middle Albian	Griman Creek Formation	Australia	A meiolaniid-like turtle. The type species is Spoochelys ormondea.
Testudo oughlamensis^[99]	Sp. nov	Valid	Gmira et al.	Pliocene		Morocco	A tortoise, a species of Testudo.
Trapalcochelys^[116]	Gen. et sp. nov	Valid	Sterli, de la Fuente & Cerda	Late Cretaceous (late Campanian to early Maastrichtian)	Allen Formation	Argentina	A relative of meiolaniids. The type species is Trapalcochelys sulcata.
Tullochelys^[94]	Gen. et sp. nov	Valid	Hutchison	Maastrichtian to early Paleocene (Puercan)	Fort Union Formation Hell Creek Formation	United States	A chelydrid. The type species is Tullochelys montanus.
Xenochelys floridensis^[117]	Sp. nov	Valid	Bourque	Late Oligocene (early Arikareean)		United States	A kinosternid, a species of Xenochelys.
Xinjiangchelys radiplicatoides^[118]	Sp. nov	Valid	Brinkman et al.	Late Middle or early Late Jurassic	Shishugou Formation	China	A species of Xinjiangchelys.
Xinjiangchelys wusu^[119]	Sp. nov	Valid	Rabi et al.	Middle Jurassic	?Qigu Formation	China	A species of Xinjiangchelys.

Thalattosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Concavispina^[120]	Gen. et sp. nov	Valid	Zhao et al.	Triassic	Xiaowa Formation	China	A thalattosaurid thalattosaur. The type species is Concavispina biseridens

Ichthyopterygians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Chaohusaurus zhangjiawanensis^[121]	Sp. nov	Valid	Chen et al.	Early Triassic	Jialingjiang Formation	China	A species of Chaohusaurus.
Gulosaurus^[122]	Gen. et sp. nov	Valid	Cuthbertson, Russell & Anderson	Early Triassic	Sulphur Mountain Formation	Canada	A grippidian. The type species is Gulosaurus helmi.
Malawania^[123]	Gen. et sp. nov	Valid	Fischer et al.	Early Cretaceous (late Hauterivian to Barremian)		Iraq	A basal member of Thunnosauria. The type species is Malawania anachronus.
Thalattoarchon^[124]	Gen. et sp. nov	Valid	Fröbisch et al.	Middle Triassic		United States	A merriamosaur ichthyosaur. The type species is Thalattoarchon saurophagis

Lepidosauromorphs

Newly named sauropterygians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Cryonectes^[125]	Gen. et sp. nov	Valid	Vincent, Bardet & Mattioli	Pliensbachian		France	A pliosaurid. The type species is Cryonectes neustriacus
Gronausaurus^[126]	Gen. et sp. nov	Valid	Hampe	Early Cretaceous (Berriasian)		Germany	A leptocleidid^[126] or an elasmosaurid.^[127] The type species is Gronausaurus wegneri
Megacephalosaurus^[128]	Gen. et sp. nov	Valid	Schumacher, Carpenter & Everhart	Late Cretaceous (middle Turonian)		United States	A pliosaurid. The type species is Megacephalosaurus eulerti.
Palatodonta^[129]	Gen. et sp. nov	Valid	Neenan, Klein & Scheyer	Early Middle Triassic		Netherlands	A relative of placodonts. The type species is Palatodonta bleekeri.
Pliosaurus carpenteri^[127]	Sp. nov	Valid	Benson et al.	Late Jurassic	Kimmeridge Clay Formation	United Kingdom	A pliosaurid, a species of Pliosaurus
Pliosaurus kevani^[127]	Sp. nov	Valid	Benson et al.	Late Jurassic	Kimmeridge Clay Formation	United Kingdom	A pliosaurid, a species of Pliosaurus
Pliosaurus westburyensis^[127]	Sp. nov	Valid	Benson et al.	Late Jurassic	Kimmeridge Clay Formation	United Kingdom	A pliosaurid, a species of Pliosaurus
Serpianosaurus germanicus^[130]	Sp. nov	Valid	Diedrich	Middle Triassic (Anisian)		Germany	A pachypleurosaur, a species of Serpianosaurus.

Newly named rhynchocephalians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Sphenotitan^[131]	Gen. et sp. nov	Valid	Martínez et al.	Late Triassic	Quebrada del Barro Formation	Argentina	An opisthodontian rhynchocephalian. The type species is Sphenotitan leyesi.

Newly named lizards

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Apsgnathus^[132]	Gen. et sp. nov	Valid	Nydam, Rowe & Cifelli	Late Cretaceous (late Campanian, Judithian)	Aguja Formation	United States	A member of Scincomorpha of uncertain phylogenetic placement. The type species is Apsgnathus triptodon.
Arcanosaurus^[133]	Gen. et sp. nov	Valid	Houssaye et al.	Early Cretaceous (Barremian or Aptian)	Cameros Basin	Spain	An anguimorph lizard, possibly a varanoid. The type species is Arcanosaurus ibericus.
Arpadosaurus sepulchralis^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Glyptosaurinae, a species of Arpadosaurus.
Barbaturex^[135]	Gen. et sp. nov	Valid	Head et al.	Late middle Eocene (37.2 ± 1.3 Ma)	Pondaung Formation	Myanmar	An iguanian lizard related to the genus Uromastyx. The type species is Barbaturex morrisoni.
Bicuspidon smikros^[136]	Sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A member of Polyglyphanodontinae, a species of Bicuspidon.
Blanosaurus^[137]	Gen. et sp. nov	Valid	Folie, Smith & Smith	Early Eocene		Belgium France	An amphisbaenian. The type species is Blanosaurus primeocaenus.
Camptognathosaurus^[137]	Gen. et sp. nov	Valid	Folie, Smith & Smith	Late Paleocene		France	An amphisbaenian. The type species is Camptognathosaurus parisiensis.
Catactegenys^[132]	Gen. et sp. nov	Valid	Nydam, Rowe & Cifelli	Late Cretaceous (late Campanian, Judithian)	Aguja Formation	United States	A night lizard. The type species is Catactegenys solaster.
Cemeterius^[138]^[139]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A platynotan lizard of uncertain phylogenetic placement. The type species is Cemeterius monstrosus.
Cnodontosaurus^[136]	Gen. et sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A member of Anguimorpha related to xenosaurids. The type species is Cnodontosaurus suchockii.
Dakotasaurus^[136]	Gen. et sp. nov	Junior homonym	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.^[136] The type species is Dakotasaurus gillettorum. The generic name turned out to be preoccupied by Dakotasaurus Branson & Mehl (1932); the lizard genus was subsequently renamed Dakotaseps.^[140]
Desertiguana^[141]	Gen. et sp. nov	Valid	Alifanov	Late Cretaceous		Mongolia	A phrynosomatid lizard. The type species is Desertiguana gobiensis.
Distortodon^[142]	Gen. et sp. nov	Valid	Makádi	Late Cretaceous (Santonian)		Hungary	A member of Polyglyphanodontinae. The type species is Distortodon rhomboideus.
Entomophontes^[134]	Gen. et 2 sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A relative of species belonging to the genus Xenosaurus. Genus contains two species: Entomophontes incrustatus and Entomophontes hutchisoni.
Eonatator coellensis^[143]	Sp. nov	Valid	Páramo	Campanian		Colombia	A mosasaur, a new species of Eonatator.
Glyptosaurus agmodon^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Glyptosaurinae, a species of Glyptosaurus.
Heterodontagama^[144]	Gen. et sp. nov	Valid	Rana et al.	Early Eocene (early to middle Ypresian)	Cambay Shale Formation	India	A member of Priscagamidae. The type species is Heterodontagama borsukae.
Indiagama^[144]	Gen. et sp. nov	Valid	Rana et al.	Early Eocene (early to middle Ypresian)	Cambay Shale Formation	India	A member of Agamidae. The type species is Indiagama gujarata.
Lamiasaura^[138]^[139]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A squamate of uncertain phylogenetic placement. The type species is Lamiasaura ferox.
Lonchisaurus^[138]^[139]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A scincomorph lizard of uncertain phylogenetic placement. The type species is Lonchisaurus trichurus.
Monocnemodon^[136]	Gen. et sp. nov	Valid	Nydam	Late Cretaceous (Santonian)	Straight Cliffs Formation	United States	A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.^[136] The type species is Monocnemodon syphakos.
Obamadon^[138]^[139]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Hell Creek Formation Lance Formation	United States	A polyglyphanodontian lizard of uncertain phylogenetic placement. The type species is Obamadon gracilis.
Odaxosaurus roosevelti^[136]	Sp. nov	Valid	Nydam	Late Cretaceous	Kaiparowits Formation	United States	A member of Anguidae, a species of Odaxosaurus.
Palaeoxantusia amara^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A night lizard, a species of Palaeoxantusia.
Palepidophyma^[134]	Gen. et 2 sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A night lizard. Genus contains two species: Palepidophyma paradisa and Palepidophyma lilliputiana.
Parasaniwa cynochoros^[136]	Sp. nov	Valid	Nydam	Late Cretaceous	Kaiparowits Formation	United States	A member of Platynota of uncertain phylogenetic placement, a species of Parasaniwa.
Pariguana^[138]^[139]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	An iguanid lizard of uncertain phylogenetic placement. The type species is Pariguana lancensis.
Pelsochamops^[145]	Gen. et sp. nov	Valid	Makádi	Late Cretaceous (Santonian)	Csehbánya Formation	Hungary	A chamopsiid. The type species is Pelsochamops infrequens.
Plesiolacerta eratosthenesi^[146]	Sp. nov	Valid	Čerňanský & Augé	Late Oligocene		Germany	A lacertid lizard, a species of Plesiolacerta.
Polyodontobaena^[137]	Gen. et sp. nov	Valid	Folie, Smith & Smith	Early Paleocene	Hainin Formation	Belgium	An amphisbaenian. The type species is Polyodontobaena belgica.
Provaranosaurus fatuus^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A relative of the Chinese crocodile lizard, a species of Provaranosaurus.
Romeosaurus^[147]	Gen. et 2 sp. nov	Valid	Palci, Caldwell & Papazzoni	Late Cretaceous (early Turonian to early Santonian)		Italy	A mosasaur related to Russellosaurus and Yaguarasaurus. Genus contains two species: Romeosaurus fumanensis and Romeosaurus sorbinii.
Schillerosaurus^[148]	Nom. nov	Valid	Nydam, Chure & Evans	Late Jurassic	Morrison Formation	United States	A replacement name for Schilleria Evans & Chure (1999).
Scincoideus grassator^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Lacertiformes, a species of Scincoideus.
Socognathus brachyodon^[138]^[139]	Sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Lance Formation	United States	A chamopsiid polyglyphanodontian lizard, a species of Socognathus.
Suratagama^[144]	Gen. et sp. nov	Valid	Rana et al.	Early Eocene (early to middle Ypresian)	Cambay Shale Formation	India	A member of Agamidae. The type species is Suratagama neeraae.
Suzanniwana revenanta^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Pleurodonta, a species of Suzanniwana.
Tiliqua laticephala^[149]	Sp. nov	Valid	Čerňanský & Hutchinson	Pliocene		Australia	A blue-tongued skink.
Utahgenys antongai^[136]	Sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A contogeniid scincomorph, a species of Utahgenys.
Webbsaurus^[136]	Gen. et sp. nov	Valid	Nydam	Late Cretaceous (Cenomanian)	Dakota Formation	United States	A scincomorph related to skinks. The type species is Webbsaurus lofgreni.
Xestops savagei^[134]	Sp. nov	Valid	Smith & Gauthier	Early Eocene	Wasatch Formation	United States	A member of Glyptosaurinae, a species of Xestops.

Newly named snakes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Cerberophis^[138]^[139]	Gen. et sp. nov	Valid	Longrich, Bhullar & Gauthier	Late Cretaceous (late Maastrichtian)	Hell Creek Formation	United States	An alethinophidian snake of uncertain phylogenetic placement. The type species is Cerberophis robustus.
Kataria^[150]	Gen. et sp. nov	Valid	Scanferla et al.	Early Paleocene (Danian)	Santa Lucía Formation	Bolivia	A macrostomatan snake related to caenophidians. The type species is Kataria anisodonta.
Nidophis^[151]	Gen. et sp. nov	Valid	Vasile, Csiki-Sava & Venczel	Late Cretaceous (Maastrichtian)	Haţeg Basin	Romania	A madtsoiid. The type species is Nidophis insularis.

Archosauromorphs

Newly named basal archosauromorphs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Ankylosuchus^[152]	Gen. et sp. nov	Valid	Lucas, Spielmann & Hunt	Late Triassic	Colorado City Formation	United States	A doswelliid. The type species is Ankylosuchus chinlegroupensis.
Asperoris^[153]	Gen. et sp. nov	Valid	Nesbitt, Butler & Gower	Middle Triassic (probably late Anisian)	Manda beds	Tanzania	A non-archosaurian archosauriform of uncertain phylogenetic placement. The type species is Asperoris mnyama.
Fuyuansaurus^[154]	Gen. et sp. nov	Valid	Fraser, Rieppel & Chun	Middle Triassic (Ladinian)	Falang Formation	China	A protorosaur. The type species is Fuyuansaurus acutirostris.
Jaxtasuchus^[155]	Gen. et sp. nov	Valid^[156]	Schoch & Sues	Middle Triassic (Ladinian)	Erfurt Formation	Germany	A doswelliid. The type species is Jaxtasuchus salomoni.
Machaeroprosopus lottorum^[157]	Sp. nov	Valid	Hungerbühler et al.	Late Triassic	Cooper Canyon Formation	United States	A phytosaur, a species of Machaeroprosopus.
Macrocnemus obristi^[158]	Sp. nov	Valid	Fraser & Furrer	Middle Triassic (Ladinian)	Prosanto Formation	Switzerland	A protorosaur, a species of Macrocnemus.
Wannia^[159]	Gen. et comb. nov	Valid	Stocker	Late Triassic		United States	A phytosaur; a new genus for "Paleorhinus" scurriensis Langston (1949).

Archosaurs

Main article: 2013 in archosaur paleontology

Other reptiles

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Dolerosaurus^[160]	Gen. et comb. nov	Valid	Butler	Late Triassic (late Carnian)	Lunz Formation	Austria	A diapsid of uncertain phylogenetic placement, a new genus for "Francosuchus" trauthi Huene (1939).
Ruhuhuaria^[161]	Gen. et sp. nov	Valid	Tsuji, Sobral & Müller	Middle Triassic	Manda Formation	Tanzania	A procolophonoid parareptile, possibly an owenettid. The type species is Ruhuhuaria reiszi.

Synapsids

Non-mammalian synapsids

Research

The postcranial skeleton of therocephalian Ictidosuchoides is described by Heidi Fourie (2013).^[162]

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Diegocanis^[163]	Gen. et sp. nov	Valid	Martínez, Fernandez & Alcober	Late Triassic	Ischigualasto Formation	Argentina	A member of Eucynodontia closely related to Ecteninion and Trucidocynodon. The type species is Diegocanis elegans.
Mandagomphodon^[164]	Gen. et comb. nov	Valid^[165]	Hopson	Middle Triassic	Manda Formation	Tanzania	A traversodontid cynodont, a new genus for "Scalenodon" hirschsoni Crompton (1972).
Niassodon^[166]	Gen. et sp. nov	Valid	Castanhinha et al.	Late Permian		Mozambique	A kingoriid emydopoid dicynodont. The type species is Niassodon mfumukasi.
Tambacarnifex^[167]	Gen. et sp. nov	Valid^[165]	Berman et al.	Early Permian	Tambach Formation	Germany	A varanodontine varanopid. The type species is Tambacarnifex unguifalcatus.

Mammals

Main article: 2013 in mammal paleontology

Other animals

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Alveopora jessicae^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Alveopora.
Alveopora oliveri^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Alveopora.
Anomalocaris kunmingensis^[169]	Sp. nov	Valid	Wang, Huang & Hu	Cambrian	Wulongqing Formation	China	A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods), a species of Anomalocaris.
Astreopora wallaceae^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Astreopora.
Aulastraea crassicalix^[170]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Aulastraea.
Cambrocornulitus^[171]	Gen. et sp. nov	Valid	Yang et al.	Early Cambrian	Qiongzhusi Formation	China	An animal of uncertain phylogenetic placement, possibly a member of Cornulitida. The type species is Cambrocornulitus rarus.
Caulastraea cummingae^[168]	Sp. nov	Valid	Bromfield	Miocene		Fiji Indonesia	A stony coral, a species of Caulastraea.
Caulastraea geoffreyi^[168]	Sp. nov	Valid	Bromfield	Miocene		Indonesia	A stony coral, a species of Caulastraea.
Conularia porcella^[172]	Sp. nov	Valid	Robson & Young	Late Ordovician (middle Katian)	Red River Formation	Canada	A conulariid, a species of Conularia.
Cyphastrea aliceae^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral belonging to the family Faviidae.
Cyphastrea buddae^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral belonging to the family Faviidae.
Cyphastrea spinea^[168]	Sp. nov	Valid	Bromfield	Miocene		Fiji	A stony coral belonging to the family Faviidae.
Euphyllia romani^[168]	Sp. nov	Valid	Bromfield	Miocene		Indonesia	A stony coral, a species of Euphyllia.
Galaxea kabairaensis^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Galaxea.
Galaxea salayarensis^[168]	Sp. nov	Valid	Bromfield	Pliocene		Indonesia	A stony coral, a species of Galaxea.
Gracilitheca astronauta^[173]	Sp. nov	Valid	Valent & Fatka	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.
Gracilitheca mirabilis^[174]	Sp. nov	Valid	Valent, Fatka & Marek	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.
Gracilitheca triangularis^[174]	Sp. nov	Valid	Valent, Fatka & Marek	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha, a species of Gracilitheca.
Heteropistophyllum^[170]	Gen. et comb. nov	Valid	Löser, Castro & Nieto	Early Cretaceous		Slovenia Spain	A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. A new genus for "Pseudopistophyllum" quinqueseptatum Turnšek & Buser (1976).
Leptoria hayae^[168]	Sp. nov	Valid	Bromfield	Miocene		Fiji	A stony coral, a species of Leptoria.
Leptoria traba^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Leptoria.
Leptoseris lauensis^[168]	Sp. nov	Valid	Bromfield	Miocene	Lau Group	Fiji	A stony coral belonging to the family Agariciidae.
Madracis crassisepta^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral belonging to the family Pocilloporidae.
Madracis labrum^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral belonging to the family Pocilloporidae.
Metaulastraea elae^[170]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Metaulastraea.
Microconchus utahensis^[175]	Sp. nov	Valid	Zatoń, Taylor & Vinn	Early Triassic	Virgin Formation	United States	A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Microconchus.
Mitrodendron ultimus^[170]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Mitrodendron.
Montipora pandolfii^[168]	Sp. nov	Valid	Bromfield	Miocene		Indonesia	A stony coral, a species of Montipora.
Montipora teresea^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Montipora.
Montipora wilsonae^[168]	Sp. nov	Valid	Bromfield	Early Pleistocene		Indonesia	A stony coral, a species of Montipora.
Nephrotheca betula^[174]	Sp. nov	Valid	Valent, Fatka & Marek	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha, a species of Nephrotheca.
Nephrotheca sophia^[173]	Sp. nov	Valid	Valent & Fatka	Cambrian	Buchava Formation	Czech Republic	A member of Hyolitha, a species of Nephrotheca.
Paraacanthogyra hispanica^[170]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Paraacanthogyra.
Paraconularia bachmanni^[176]	Sp. nov	Valid	Barth et al.	Late Triassic		Germany	A conulariid, a species of Paraconularia.
Paranomalocaris^[169]	Gen. et sp. nov	Valid	Wang, Huang & Hu	Cambrian	Wulongqing Formation	China	A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods). The type species is Paranomalocaris multisegmentalis.
Platygyra renemai^[168]	Sp. nov	Valid	Bromfield	Miocene		Papua New Guinea	A stony coral, a species of Platygyra.
Pleurodendron^[170]	Gen. et sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. The type species is Pleurodendron prebeticum.
Quasiaulacera^[177]	Gen. et 2 sp. nov	Valid	Copper, Stock & Jin	Late Ordovician (Hirnantian)	Ellis Bay Formation	Canada	A stromatoporoid. Genus contains two species: Quasiaulacera stellata and Q. occidua.
Silingastraea valentina^[170]	Sp. nov	Valid	Löser, Castro & Nieto	Early Cretaceous (Albian)		Spain	A stony coral, a species of Silingastraea.
Spinitheca^[178]	Gen. et sp. nov	Valid	Novozhilova	Early Cambrian (Atdabanian)		Russia	A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is Spinitheca sysoievi.
Symphyllia densium^[168]	Sp. nov	Valid	Bromfield	Early Pleistocene		Indonesia	A stony coral belonging to the family Mussidae.
Turbinaria jacksoni^[168]	Sp. nov	Valid	Bromfield	Miocene		Fiji	A stony coral, a species of Turbinaria.

Other organisms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Honeggeriella^[179]	Gen. et sp. nov	Valid	Matsunaga, Stockey & Tomescu	Early Cretaceous (Valanginian-Hauterivian boundary)	Longarm Formation	Canada	A heteromerous lichen. The type species is Honeggeriella complexa.
Plumalina brevis^[180]	Sp. nov	Valid	Muscente & Allmon	Devonian (Frasnian)	Ithaca Formation	United States	An organism of uncertain phylogenetic placement, argued by different authors to be a hydrozoan^[180] or a Caulerpa-like alga.^[181]
Plumalina tenera^[180]	Sp. nov	Valid	Muscente & Allmon	Silurian (Wenlock)	Rochester Shale	United States	An organism of uncertain phylogenetic placement, argued by different authors to be a hydrozoan^[180] or a Caulerpa-like alga.^[181]

References

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↑ Richard J. Aldridge; Duncan J. E. Murdock; Sarah E. Gabbott; Johannes N. Theron (2013). "A 17-element conodont apparatus from the Soom Shale Lagerstätte (Upper Ordovician), South Africa". Palaeontology. 56 (2): 261–276. doi:10.1111/j.1475-4983.2012.01194.x.
1 2 3 4 5 6 7 8 9 10 11 12 13 Norman M. Savage (2013). Late Devonian conodonts from northwestern Thailand. Bourland Printing / Trinity Press. pp. 1–48. ISBN 978-0-578-11919-9.
↑ Hossein Gholamalian; Vachik Hairapetian; Nahid Barfehei; Soheyla Mangelian; Parvaneh Faridi (2013). "Givetian–Frasnian boundary conodonts from Kerman Province, central Iran". Rivista Italiana di Paleontologia e Stratigrafia. 119 (2): 133–146. doi:10.13130/2039-4942/6028.
↑ Gilbert Klapper; Stanislava Vodrážková (2013). "Ontogenetic and intraspecific variation in the late Emsian – Eifelian (Devonian) conodonts Polygnathus serotinus and P. bultyncki in the Prague Basin (Czech Republic) and Nevada (western U.S.)". Acta Geologica Polonica. 63 (2): 153–174. doi:10.2478/agp-2013-0006.
↑ Gustavo Gabriel Voldman; Guillermo Luis Albanesi; Fernando Javier Zeballo; Cesar Ruben Monaldi (2013). "Early Ordovician (Late Floian) conodonts from the Zenta range, Cordillera Oriental, NW Argentina". Publicación Especial - Asociación Paleontológica Argentina. 13: 123–128.
↑ Gustavo G. Voldman; Guillermo L. Albanesi; Gladys Ortega; María Eugenia Giuliano; Carlos Ruben Monaldi (2016). "New conodont taxa and biozones from the Lower Ordovician of the Cordillera Oriental, NW Argentina". Geological Journal. in press. doi:10.1002/gj.2766.
↑ Robert Holmes; David S. Berman; Jason S. Anderson (2013). "A new dissorophid (Temnospondyli, Dissorophoidea) from the Early Permian of New Mexico (United States)". Comptes Rendus Palevol. 12 (7–8): 419–435. doi:10.1016/j.crpv.2013.07.002.
↑ Tomasz Sulej; Grzegorz Niedźwiedzki (2013). "A new large capitosaur temnospondyl amphibian from the Early Triassic of Poland". Acta Palaeontologica Polonica. 58 (1): 65–75. doi:10.4202/app.2011.0025.
↑ Hillary C. Maddin; Nadia B. Fröbisch; David C. Evans; Andrew R. Milner (2013). "Reappraisal of the Early Permian amphibamid Tersomius texensis and some referred material". Comptes Rendus Palevol. 12 (7–8): 447–461. doi:10.1016/j.crpv.2013.06.007.
↑ Rainer R. Schoch; Hans-Dieter Sues (2013). "A new dissorophid temnospondyl from the Lower Permian of north-central Texas". Comptes Rendus Palevol. 12 (7–8): 437–445. doi:10.1016/j.crpv.2013.04.002.
↑ Jason S. Anderson; John R. Bolt (2013). "New information on amphibamids (Tetrapoda, Temnospondyli) from Richards Spur (Fort Sill), Oklahoma". Journal of Vertebrate Paleontology. 33 (3): 553–567. doi:10.1080/02724634.2013.726676.
↑ Andrew R. Milner; Rainer R. Schoch (2013). "Trimerorhachis (Amphibia: Temnospondyli) from the Lower Permian of Texas and New Mexico: cranial osteology, taxonomy and biostratigraphy". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 270 (1): 91–128. doi:10.1127/0077-7749/2013/0360.
↑ Sabine Glienke (2013). "A taxonomic revision of Batropetes (Amphibia, Microsauria) from the Rotliegend (basal Permian) of Germany". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 269 (1): 73–96. doi:10.1127/0077-7749/2013/0336.
1 2 Adam K. Huttenlocker; Jason D. Pardo; Bryan J. Small; Jason S. Anderson (2013). "Cranial morphology of recumbirostrans (Lepospondyli) from the Permian of Kansas and Nebraska, and early morphological evolution inferred by micro-computed tomography". Journal of Vertebrate Paleontology. 33 (3): 540–552. doi:10.1080/02724634.2013.728998.
↑ Amy C. Henrici; Ana M. Báez & Lance Grande (2013). "Aerugoamnis paulus, New Genus and New Species (Anura: Anomocoela): First Reported Anuran from the Early Eocene (Wasatchian) Fossil Butte Member of the Green River Formation, Wyoming". Annals of Carnegie Museum. 81 (4): 295–309. doi:10.2992/007.081.0402.
1 2 Annelise Folie; Rajendra S. Rana; Kenneth D. Rose; Ashok Sahni; Kishor Kumar; Lachham Singh & Thierry Smith (2013). "Early Eocene frogs from Vastan Lignite Mine, Gujarat, India". Acta Palaeontologica Polonica. 58 (3): 511–524. doi:10.4202/app.2011.0063.
1 2 Ana María Báez (2013). "Anurans from the Early Cretaceous Lagerstätte of Las Hoyas, Spain: New evidence on the Mesozoic diversification of crown-clade Anura". Cretaceous Research. 41: 90–106. doi:10.1016/j.cretres.2012.11.002.
↑ Rainer R. Schoch; Michael W. Rasser (2013). "A new salamandrid from the Miocene Randeck Maar, Germany". Journal of Vertebrate Paleontology. 33 (1): 58–66. doi:10.1080/02724634.2012.716113.
1 2 TH Worthy; AJD Tennyson; RP Scofield & SJ Hand (2013). "Early Miocene fossil frogs (Anura: Leiopelmatidae) from New Zealand". Journal of the Royal Society of New Zealand. 43 (4): 211–230. doi:10.1080/03036758.2013.825300.
↑ Liping Dong; Zbyněk Roček; Yuan Wang & Marc E H. Jones (2013). "Anurans from the Lower Cretaceous Jehol Group of Western Liaoning, China". PLoS ONE. 8 (7): e69723. doi:10.1371/journal.pone.0069723.
↑ David G. Demar Jr. (2013). "A new fossil salamander (Caudata, Proteidae) from the Upper Cretaceous (Maastrichtian) Hell Creek Formation, Montana, U.S.A.". Journal of Vertebrate Paleontology. 33 (3): 588–598. doi:10.1080/02724634.2013.734887.
↑ Davit Vasilyan; Madelaine Böhme; Viacheslav M. Chkhikvadze; Yuriy A. Semenov; Walter G. Joyce (2013). "A new giant salamander (Urodela, Pancryptobrancha) from the Miocene of Eastern Europe (Grytsiv, Ukraine)". Journal of Vertebrate Paleontology. 33 (2): 301–318. doi:10.1080/02724634.2013.722151.
↑ Steven C. Sweetman & James D. Gardner (2013). "A new albanerpetontid amphibian from the Early Cretaceous (Barremian) Wessex Formation of the Isle of Wight, southern England". Acta Palaeontologica Polonica. 58 (2): 295–324. doi:10.4202/app.2011.0109.
↑ Igor G. Danilov; Elena V. Syromyatnikova; Pavel P. Skutschas; Tatyana M. Kodrul; Jianhua Jin (2013). "The first 'true' Adocus (Testudines, Adocidae) from the Paleogene of Asia". Journal of Vertebrate Paleontology. 33 (5): 1071–1080. doi:10.1080/02724634.2013.768254.
1 2 3 4 5 J. Howard Hutchison (2013). "New turtles from the Paleogene of North America". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 477–497. doi:10.1007/978-94-007-4309-0_26. ISBN 978-94-007-4308-3.
↑ Robert E. Weems; James L. Knight (2013). "A new species of Bairdemys (Pelomedusoides: Podocnemididae) from the Oligocene (Early Chattian) Chandler Bridge Formation of South Carolina, USA, and its paleobiogeographic implications for the genus". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 289–303. doi:10.1007/978-94-007-4309-0_18. ISBN 978-94-007-4308-3.
1 2 Robert M. Sullivan; Steven E. Jasinski; Spencer G. Lucas (2013). "Re-assessment of Late Campanian (Kirtlandian) turtles from the Upper Cretaceous Fruitland and Kirtland Formations, San Juan Basin, New Mexico, USA". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 337–387. doi:10.1007/978-94-007-4309-0_20. ISBN 978-94-007-4308-3.
↑ Haiyan Tong; Peter Meylan (2013). "Morphology and relationships of Brachyopsemys tingitana gen. et sp. nov. from the Early Paleocene of Morocco and recognition of the new eucryptodiran turtle family: Sandownidae". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 187–212. doi:10.1007/978-94-007-4309-0_13. ISBN 978-94-007-4308-3.
↑ A. Pérez-García; X. Murelaga (2013). "Camerochelys vilanovai gen. et sp. nov., a new pan-cryptodiran turtle in the Early Cretaceous of the Iberian Range (Spain)". Cretaceous Research. 41: 143–149. doi:10.1016/j.cretres.2012.12.002.
1 2 Saloua Gmira; France de Lapparent de Broin; Denis Geraads; David Lefèvre; Abderrahim Mohib & Jean-Paul Raynal (2013). "Les tortues du Pliocène d'Ahl al Oughlam (Casablanca, Maroc) et de localités mio-pliocènes avoisinantes". Geodiversitas. 35 (3): 691–733. doi:10.5252/g2013n3a9.
↑ Donald B. Brinkman; Chong-Xi Yuan; Qiang Ji; Da-Qing Li; Hai-Lu You (2013). "A new turtle from the Xiagou Formation (Early Cretaceous) of Changma Basin, Gansu Province, P. R. China". Palaeobiodiversity and Palaeoenvironments. 93 (3): 367–382. doi:10.1007/s12549-013-0113-0.
↑ Adam M. Yates (2013). "A new species of long-necked turtle (Pleurodira: Chelidae: Chelodina) from the late Miocene Alcoota Local Fauna, Northern Territory, Australia". PeerJ. 1: e170. doi:10.7717/peerj.170.
↑ Gerardo Gabriel Zacarías; Marcelo Saúl de la Fuente; Marta Susana Fernández; Alfredo Eduardo Zurita (2013). "Nueva especie de tortuga terrestre gigante del género Chelonoidis Fitzinger, 1835 (Cryptodira: Testudinidae), del miembro inferior de la Formación Toropí/ Yupoí (Pleistoceno tardío/ Lujanense), Bella Vista, Corrientes, Argentina". Ameghiniana. 50 (3): 298–318. doi:10.5710/amgh.05.03.2013.549.
1 2 Robert E. Weems; Reed A. George (2013). "Amphibians and nonmarine turtles from the Miocene Calvert Formation of Delaware, Maryland, and Virginia (USA)". Journal of Paleontology. 87 (4): 570–588. doi:10.1666/12-071.
↑ Wilailuck Naksri; Haiyan Tong; Komsorn Lauprasert; Varavudh Suteethorn; Julien Claude (2013). "A new species of Cuora (Testudines: Geoemydidae) from the Miocene of Thailand and its evolutionary significance". Geological Magazine. 150 (5): 908–922. doi:10.1017/S0016756812001082.
↑ Ren Hirayama; Shinji Isaji; Tsuyoshi Hibino (2013). "Kappachelys okurai gen. et sp. nov., a new stem soft-shelled turtle from the Early Cretaceous of Japan". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 179–185. doi:10.1007/978-94-007-4309-0_12. ISBN 978-94-007-4308-3.
↑ Paul C. Sereno; Sara J. ElShafie (2013). "A new long-necked turtle, Laganemys tenerensis (Pleurodira: Araripemydidae), from the Elrhaz Formation (Aptian–Albian) of Niger". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 215–250. doi:10.1007/978-94-007-4309-0_14. ISBN 978-94-007-4308-3.
↑ Adán Pérez-García; France de Lapparent de Broin (2013). "A new species of Neochelys (Chelonii, Podocnemididae) from the Ypresian (Early Eocene) of the South of France". Comptes Rendus Palevol. 12 (5): 269–277. doi:10.1016/j.crpv.2013.05.011.
↑ Derek W. Larson; Nicholas R. Longrich; David C. Evans; Michael J. Ryan (2013). "A new species of Neurankylus from the Milk River Formation (Cretaceous: Santonian) of Alberta, Canada, and a revision of the type species N. eximius". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 389–405. doi:10.1007/978-94-007-4309-0_21. ISBN 978-94-007-4308-3.
↑ Georgios L. Georgalis; Evangelos Velitzelos; Dimitrios E. Velitzelos; Benjamin P. Kear (2013). "Nostimochelone lampra gen. et sp. nov., an enigmatic new podocnemidoidean turtle from the Early Miocene of northern Greece". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 277–287. doi:10.1007/978-94-007-4309-0_17. ISBN 978-94-007-4308-3.
↑ Akio Takahashi; Kimihiko Ōki; Takahiro Ishido; Ren Hirayama (2013). "A new species of the genus Ocadia (Testudines: Geoemydidae) from the middle Miocene of Tanegashima Island, southwestern Japan and its paleogeographic implications". Zootaxa. 3647 (4): 527–540. doi:10.11646/zootaxa.3647.4.3.
↑ Nathalie Bardet; Nour-Eddine Jalil; France de Lapparent de Broin; Damien Germain; Olivier Lambert & Mbarek Amaghzaz (2013). "A Giant Chelonioid Turtle from the Late Cretaceous of Morocco with a Suction Feeding Apparatus Unique among Tetrapods". PLoS ONE. 8 (7): e63586. doi:10.1371/journal.pone.0063586.
↑ I. G. Danilov; V. B. Sukhanov (2013). "A new basal testudinoid turtle (Testudinoidea: "Lindholmemydidae") from the Upper Paleocene of Mongolia". Paleontological Journal. 47 (1): 104–113. doi:10.1134/S0031030113010036.
↑ Lu Li; Haiyan Tong; Kebai Wang; Shuqing Chen; Xing Xu (2013). "Lindholmemydid turtles (Cryptodira: Testudinoidea) from the Late Cretaceous of Shandong Province, China". Annales de Paléontologie. 99 (3): 243–259. doi:10.1016/j.annpal.2013.07.003.
↑ Haiyan Tong; Donald Brinkman (2013). "A new species of Sinemys (Testudines: Cryptodira: Sinemydidae) from the Early Cretaceous of Inner Mongolia, China". Palaeobiodiversity and Palaeoenvironments. 93 (3): 355–366. doi:10.1007/s12549-012-0110-8.
↑ Elizabeth T. Smith; Benjamin P. Kear (2013). "Spoochelys ormondea gen. et sp. nov., an Archaic Meiolaniid-Like Turtle from the Early Cretaceous of Lightning Ridge, Australia". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 121–146. doi:10.1007/978-94-007-4309-0_9. ISBN 978-94-007-4308-3.
↑ Juliana Sterli; Marcelo S. de la Fuente; Ignacio A. Cerda (2013). "A new species of meiolaniform turtle and a revision of the Late Cretaceous Meiolaniformes of South America". Ameghiniana. 50 (2): 240–256. doi:10.5710/amgh.16.01.2013.582.
↑ Jason R. Bourque (2013). "Fossil Kinosternidae from the Oligocene and Miocene of Florida, USA". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 459–475. doi:10.1007/978-94-007-4309-0_25. ISBN 978-94-007-4308-3.
↑ Donald B. Brinkman; David A. Eberth; Xing Xu; James M. Clark; Xiao-Chun Wu (2013). "Turtles from the Jurassic Shishugou Formation of the Junggar Basin, People's Republic of China, with comments on the basicranial region of basal eucryptodires". In Donald B. Brinkman; Patricia A. Holroyd; James D. Gardner. Morphology and Evolution of Turtles. Springer. pp. 147–172. doi:10.1007/978-94-007-4309-0_10. ISBN 978-94-007-4308-3.
↑ Márton Rabi; Chang-Fu Zhou; Oliver Wings; Sun Ge; Walter G. Joyce (2013). "A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles". BMC Evolutionary Biology. 13: Article number 203. doi:10.1186/1471-2148-13-203.
↑ Zhao Li-Jun; Liu Jun; Li Chun; He Tao (2013). "A new thalattosaur, Concavispina biseridens gen. et sp. nov. from Guanling, Guizhou, China" (PDF). Vertebrata PalAsiatica. 51 (1): 24–28.
↑ Xiaohong Chen; P. Martin Sander; Long Cheng; Xiaofeng Wang (2013). "A New Triassic Primitive Ichthyosaur from Yuanan, South China". Acta Geologica Sinica (English Edition). 87 (3): 672–677. doi:10.1111/1755-6724.12078.
↑ Robin S. Cuthbertson; Anthony P. Russell & Jason S. Anderson (2013). "Cranial morphology and relationships of a new grippidian (Ichthyopterygia) from the Vega-Phroso Siltstone Member (Lower Triassic) of British Columbia, Canada". Journal of Vertebrate Paleontology. 33 (4): 831–847. doi:10.1080/02724634.2013.755989.
↑ Valentin Fischer; Robert M. Appleby; Darren Naish; Jeff Liston; James B. Riding; Stephen Brindley; Pascal Godefroit (2013). "A basal thunnosaurian from Iraq reveals disparate phylogenetic origins for Cretaceous ichthyosaurs". Biology Letters. 9 (4): 20130021. doi:10.1098/rsbl.2013.0021.
↑ Nadia B. Fröbisch; Jörg Fröbisch; P. Martin Sander; Lars Schmitz & Olivier Rieppel (2013). "Macropredatory ichthyosaur from the Middle Triassic and the origin of modern trophic networks". Proceedings of the National Academy of Sciences of the United States of America. 110 (4): 1393–1397. doi:10.1073/pnas.1216750110.
↑ Peggy Vincent; Nathalie Bardet & Emanuela Mattioli (2013). "A new pliosaurid from the Pliensbachian (Early Jurassic) of Normandy (Northern France)". Acta Palaeontologica Polonica. 58 (3): 471–485. doi:10.4202/app.2011.0113.
1 2 Oliver Hampe (2013). "The forgotten remains of a leptocleidid plesiosaur (Sauropterygia: Plesiosauroidea) from the Early Cretaceous of Gronau (Münsterland, Westphalia, Germany)". Paläontologische Zeitschrift. 87 (4): 473–491. doi:10.1007/s12542-013-0175-3.
1 2 3 4 Roger B. J. Benson; Mark Evans; Adam S. Smith; Judyth Sassoon; Scott Moore-Faye; Hilary F. Ketchum & Richard Forrest (2013). "A Giant Pliosaurid Skull from the Late Jurassic of England". PLoS ONE. 8 (5): e65989. doi:10.1371/journal.pone.0065989.
↑ Bruce A. Schumacher; Kenneth Carpenter; Michael J. Everhart (2013). "A new Cretaceous Pliosaurid (Reptilia, Plesiosauria) from the Carlile Shale (middle Turonian) of Russell County, Kansas". Journal of Vertebrate Paleontology. 33 (3): 613–628. doi:10.1080/02724634.2013.722576.
↑ James M. Neenan; Nicole Klein & Torsten M. Scheyer (2013). "European origin of placodont marine reptiles and the evolution of crushing dentition in Placodontia". Nature Communications. 4: 1621. doi:10.1038/ncomms2633.
↑ Cajus G. Diedrich (2013). "The marine pachypleurosaur Serpianosaurus germanicus nov. spec. – skeleton and isolated bone remains from the Pelsonian (Middle Triassic) of the European Germanic basin carbonate intertidals and its paleobiology and taphonomy" (PDF). New Mexico Museum of Natural History and Science Bulletin. 61: 159–168.
↑ Ricardo N. Martínez; Cecilia Apaldetti; Carina E. Colombi; Angel Praderio; Eliana Fernandez; Paula Santi Malnis; Gustavo A. Correa; Diego Abelin & Oscar Alcober (2013). "A new sphenodontian (Lepidosauria: Rhynchocephalia) from the Late Triassic of Argentina and the early origin of the herbivore opisthodontians". Proceedings of the Royal Society B. 280 (1772): 20132057. doi:10.1098/rspb.2013.2057.
1 2 Randall L. Nydam; Timothy B. Rowe; Richard L. Cifelli (2013). "Lizards and snakes of the Terlingua Local Fauna (late Campanian), Aguja Formation, Texas, with comments on the distribution of paracontemporaneous squamates throughout the Western Interior of North America". Journal of Vertebrate Paleontology. 33 (5): 1081–1099. doi:10.1080/02724634.2013.760467.
↑ Alexandra Houssaye; Jean-Claude Rage; Fidel Torcida Fernández-Baldor; Pedro Huerta; Nathalie Bardet; Xabier Pereda Suberbiola (2013). "A new varanoid squamate from the Early Cretaceous (Barremian–Aptian) of Burgos, Spain". Cretaceous Research. 41: 127–135. doi:10.1016/j.cretres.2012.11.005.
1 2 3 4 5 6 7 8 9 Krister T. Smith; Jacques A. Gauthier (2013). "Early Eocene Lizards of the Wasatch Formation near Bitter Creek, Wyoming: Diversity and Paleoenvironment during an Interval of Global Warming". Bulletin of the Peabody Museum of Natural History. 54 (2): 135–230. doi:10.3374/014.054.0205.
↑ Jason J. Head; Gregg F. Gunnell; Patricia A. Holroyd; J. Howard Hutchison; Russell L. Ciochon (2013). "Giant lizards occupied herbivorous mammalian ecospace during the Paleogene greenhouse in Southeast Asia". Proceedings of the Royal Society B. 280 (1763): 20130665. doi:10.1098/rspb.2013.0665.
1 2 3 4 5 6 7 8 9 10 Randall L. Nydam (2013). "Lizards and Snakes from the Cenomanian through Campanian of Southern Utah: Filling the Gap in the Fossil Record of Squamata from the Late Cretaceous of the Western Interior of North America". In Alan L. Titus; Mark A. Loewen. At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah. Indiana University Press. pp. 370–423. ISBN 978-0-253-00896-1.
1 2 3 Annelise Folie; Richard Smith; Thierry Smith (2013). "New amphisbaenian lizards from the Early Paleogene of Europe and their implications for the early evolution of modern amphisbaenians". Geologica Belgica. 16 (4): 227–235.
1 2 3 4 5 6 7 Nicholas R. Longrich; Bhart-Anjan S. Bhullar; Jacques A. Gauthier (2012). "Mass extinction of lizards and snakes at the Cretaceous–Paleogene boundary". Proceedings of the National Academy of Sciences of the United States of America. 109 (52): 21396–21401. doi:10.1073/pnas.1211526110. PMC 3535637. PMID 23236177.
1 2 3 4 5 6 7 Nicholas R. Longrich; Bhart-Anjan S. Bhullar; Jacques A. Gauthier (2013). "Correction for "Mass extinction of lizards and snakes at the Cretaceous–Paleogene boundary," by Nicholas R. Longrich, Bhart-Anjan S. Bhullar, and Jacques A. Gauthier, which appeared in issue 52, December 26, 2012, of Proc Natl Acad Sci USA (109:21396–21401; first published December 10, 2012; 10.1073/pnas.1211526110)". Proceedings of the National Academy of Sciences of the United States of America. 110 (16): 6608. doi:10.1073/pnas.1303907110.
↑ Randall L. Nydam (2014). "Dakotaseps gen. nov., a replacement name for the lizard genus Dakotasaurus Nydam 2013, a junior homonym of the ichnotaxon Dakotasaurus Branson and Mehl 1932". Zootaxa. 3900 (1): 150. doi:10.11646/zootaxa.3900.1.11.
↑ V. R. Alifanov (2013). "Desertiguana gobiensis gen. et sp. nov., a new lizard (Phrynosomatidae, Iguanomorpha) from the Upper Cretaceous of Mongolia". Paleontological Journal. 47 (4): 417–424. doi:10.1134/S0031030113040023.
↑ László Makádi (2013). "A new polyglyphanodontine lizard (Squamata: Borioteiioidea) from the Late Cretaceous Iharkút locality (Santonian, Hungary)". Cretaceous Research. 46: 166–176. doi:10.1016/j.cretres.2013.08.001.
↑ Páramo-Fonseca, M. E. (2013). "Eonatator coellensis nov. sp. (Squamata: Mosasauridae), nueva especie del Cretácico Superior de Colombia" [Eonatator coellensis nov. sp. (Squamata: Mosasauridae), a new species from the Upper Cretaceous of Colombia]. Revista de la Academia Colombiana de Ciencias (in Spanish). 37 (145): 499–518. ISSN 0370-3908.
1 2 3 Rajendra S. Rana; Marc Augé; Annelise Folie; Kenneth D. Rose; Kishor Kumar; Lachham Singh; Ashok Sahni; Thierry Smith (2013). "High diversity of acrodontan lizards in the Early Eocene Vastan Lignite Mine of India". Geologica Belgica. 16 (4): 290–301.
↑ László Makádi (2013). "The first known chamopsiid lizard (Squamata) from the Upper Cretaceous of Europe (Csehbánya Formation; Hungary, Bakony Mts)". Annales de Paléontologie. 99 (3): 261–274. doi:10.1016/j.annpal.2013.07.002.
↑ Andrej Čerňanský; Marc Louis Augé (2013). "New species of the genus Plesiolacerta (Squamata: Lacertidae) from the upper Oligocene (MP28) of Southern Germany and a revision of the type species Plesiolacerta lydekkeri". Palaeontology. 56 (1): 79–94. doi:10.1111/j.1475-4983.2012.01167.x.
↑ Alessandro Palci; Michael W. Caldwell; Cesare A. Papazzoni (2013). "A new genus and subfamily of mosasaurs from the Upper Cretaceous of northern Italy". Journal of Vertebrate Paleontology. 33 (3): 599–612. doi:10.1080/02724634.2013.731024.
↑ Randall L. Nydam; Daniel J. Chure; Susan E. Evans (2013). "Schillerosaurus gen. nov., a replacement name for the lizard genus Schilleria Evans and Chure, 1999 a junior homonym of Schilleria Dahl, 1907" (PDF). Zootaxa. 3736 (1): 99–100. doi:10.11646/zootaxa.3736.1.6.
↑ Andrej Čerňanský; Mark N. Hutchinson (2013). "A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales, Australia)". Alcheringa: An Australasian Journal of Palaeontology. 37 (1): 131–136. doi:10.1080/03115518.2012.715326.
↑ Agustín Scanferla; Hussam Zaher; Fernando E. Novas; Christian de Muizon; Ricardo Céspedes (2013). "A New Snake Skull from the Paleocene of Bolivia Sheds Light on the Evolution of Macrostomatans". PLoS ONE. 8 (3): e57583. doi:10.1371/journal.pone.0057583.
↑ Ştefan Vasile; Zoltán Csiki-Sava; Márton Venczel (2013). "A new madtsoiid snake from the Upper Cretaceous of the Haţeg Basin, western Romania". Journal of Vertebrate Paleontology. 33 (5): 1100–1119. doi:10.1080/02724634.2013.764882.
↑ Spencer G. Lucas; Justin A. Spielmann; Adrian P. Hunt (2013). "A new doswelliid archosauromorph from the Upper Triassic of West Texas" (PDF). New Mexico Museum of Natural History and Science Bulletin. 61: 382–388.
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↑ http://www.zoobank.org/urn:lsid:zoobank.org:pub:4A400628-07F3-4D1B-9CA8-336C379721FA
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↑ Nicholas Fraser; Heinz Furrer (2013). "A new species of Macrocnemus from the Middle Triassic of the eastern Swiss Alps". Swiss Journal of Geosciences. 106 (2): 199–206. doi:10.1007/s00015-013-0137-5.
↑ Michelle R. Stocker (2013). "A new taxonomic arrangement for Paleorhinus scurriensis". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 251–263. doi:10.1017/S1755691013000340.
↑ Richard J. Butler (2013). "'Francosuchus' trauthi is not Paleorhinus: implications for Late Triassic vertebrate biostratigraphy". Journal of Vertebrate Paleontology. 33 (4): 858–864. doi:10.1080/02724634.2013.740542.
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↑ Heidi Fourie (2013). "The postcranial description of Ictidosuchoides (Therapsida: Therocephalia: Baurioidea)". Annals of the Ditsong National Museum of Natural History. 3: 1–10.
↑ Ricardo N. Martínez; Eliana Fernandez; Oscar A. Alcober (2013). "A new non-mammaliaform eucynodont from the Carnian-Norian Ischigualasto Formation, Northwestern Argentina". Revista Brasileira de Paleontologia. 16 (1): 61–76. doi:10.4072/rbp.2013.1.05.
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1 2 http://zoobank.org/References/3D07579F-44FA-46B7-9A20-D55D3B51E682
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↑ David S. Berman; Amy C. Henrici; Stuart S. Sumida; Thomas Martens; Valerie Pelletier (2013). "First European record of a varanodontine (Synapsida: Varanopidae): member of a unique Early Permian upland paleoecosystem, Tambach Basin, central Germany". In Christian F. Kammerer; Kenneth D. Angielczyk; Jörg Fröbisch. Early Evolutionary History of the Synapsida. Springer. pp. 69–86. doi:10.1007/978-94-007-6841-3_5. ISBN 978-94-007-6840-6.
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 Kate Bromfield (2013). "Neogene corals from the Indo-Pacific: Indonesia, Papua New Guinea and Fiji" (PDF). Bulletins of American Paleontology. 387: 1–127.
1 2 YuanYuan Wang; DiYing Huang; ShiXue Hu (2013). "New anomalocardid frontal appendages from the Guanshan biota, eastern Yunnan". Chinese Science Bulletin. 58 (32): 3937–3942. doi:10.1007/s11434-013-5908-x.
1 2 3 4 5 6 7 Hannes Löser; José Manuel Castro; Luis M. Nieto (2013). "Late Albian Scleractinian corals from the Prebetic Zone (SE Spain)". Palaeontographica Abteilung A. 301 (1–2): 1–62.
↑ Xianfeng Yang; Olev Vinn; Xianguang Hou; Xinglei Tian (2013). "New tubicolous problematic fossil with some "lophophorate" affinities from the Early Cambrian Chengjiang biota in south China". GFF. 135 (2): 184–190. doi:10.1080/11035897.2013.801035.
↑ Sean P. Robson; Graham A. Young (2013). "Late Ordovician conulariids from Manitoba, Canada". Journal of Paleontology. 87 (5): 775–785. doi:10.1666/12-0370.
1 2 Martin Valent; Oldřich Fatka (2013). "Gracilitheca astronauta n. sp. and Nephrotheca sophia n. sp. (Hyolitha, Orthothecida) from the Cambrian of Czech Republic". Annales de Paléontologie. 99 (3): 207–216. doi:10.1016/j.annpal.2013.03.002.
1 2 3 Martin Valent; Oldřich Fatka; Ladislav Marek (2013). "Gracilitheca and Nephrotheca (Hyolitha, Orthothecida) in the Cambrian of the Barrandian area, Czech Republic". Alcheringa: An Australasian Journal of Palaeontology. 37 (1): 115–124. doi:10.1080/03115518.2012.709446.
↑ Michał Zatoń; Paul D. Taylor; Olev Vinn (2013). "Early Triassic (Spathian) Post-Extinction Microconchids from Western Pangea". Journal of Paleontology. 87 (1): 159–165. doi:10.1666/12-060R.1.
↑ Gregor Barth; Heinz W. Kozur; Matthias Franz; Robert E. Weems (2013). "Paraconularia bachmanni n. sp. from the Germanic Upper Triassic, the first fresh-water occurrence of a conulariid" (PDF). New Mexico Museum of Natural History and Science Bulletin. 61: 42–47.
↑ Paul Copper; Carl W. Stock; Jisuo Jin (2013). "Quasiaulacera, a new Hirnantian (Late Ordovician) aulaceratid stromatoporoid genus from Anticosti Island, Canada". Journal of Paleontology. 87 (4): 664–676. doi:10.1666/12-150.
↑ N. V. Novozhilova (2013). "Spinitheca gen. nov.—A new orthothecimorph hyolith genus from the Lower Cambrian of the Siberian platform". Paleontological Journal. 47 (2): 136–138. doi:10.1134/S0031030113020081.
↑ Kelly K. S. Matsunaga; Ruth A. Stockey; Alexandru M. F. Tomescu (2013). "Honeggeriella complexa gen. et sp. nov., a heteromerous lichen from the Lower Cretaceous of Vancouver Island (British Columbia, Canada)". American Journal of Botany. 100 (2): 450–459. doi:10.3732/ajb.1200470.
1 2 3 4 A. D. Muscente; Warren D. Allmon (2013). "Revision of the hydroid Plumalina Hall, 1858 in the Silurian and Devonian of New York". Journal of Paleontology. 87 (4): 710–725. doi:10.1666/12-125.
1 2 Jerzy Dzik; Andrzej Baliński; Yuanlin Sun (2016). "An Early Ordovician clonal organism from China with a zig-zagged suture on branches". Bulletin of Geosciences. 91 (2): 319–329. doi:10.3140/bull.geosci.1575.

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