List of pterosaur genera

This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered pterosaurian. The list currently includes 213 genera.

Scope and terminology

There is no official, canonical list of pterosaur genera, but the most thorough attempts can be found at the Pterosauria section of Mikko Haaramo's Phylogeny Archive,^[1] the Genus Index at Mike Hanson's The Pterosauria,^[2] supplemented by the Pterosaur Species List,^[3] and in the fourth supplement of Donald F. Glut's Dinosaurs: The Encyclopedia series.^[4]

Authors and year

The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the description was published.

Status

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

Age

The age column denotes the epoch of geologic time to which the fossils date. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as age N/A because there was never a time in which a pterosaur by that generic name actually lived.

Location and notes

The location column designates the geographic region where remains of the relevant genus have been found. The regions used are continents except in the case of smaller landmasses (e.g. Cuba). Political bodies, being non-existent in the Mesozoic are not used to indicate genera locations. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as location N/A because there was never a place in which a pterosaur by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.

The list

Genera

Genus	Authors	Year	Status	Age	Location	Notes
Aerodactylus^[5]	Vidovic and Martill	2014	Valid	6 !Late Jurassic	Europe
Aerotitan^[6]	Novas et al.	2012	Valid.	8 !Late Cretaceous	S. America
Aetodactylus^[7]	Myers	2010	Valid.	8 !Late Cretaceous	N. America
Aidachar^[8]	Nesov	1981	Misidentification.	N/A	N/A	Actually a teleost fish whose remains were originally mistaken for jaw fragments of a ctenochasmatid. The mistake was corrected in 1986.
Alamodactylus^[9]	Andres Myers	2013	Valid.	8 !Late Cretaceous^[9]	N. America^[9]
Alanqa^[10]	Ibrahim et al.	2010	Valid	8 !Late Cretaceous	Africa
Allkaruen	Codorniú et al.	2016	Valid.	4 !Early Jurassic or 5 !Middle Jurassic	S. America
Amblydectes	Hooley	1914	Nomen dubium.	7 !Early Cretaceous	Europe
Angustinaripterus^[11]	He Xinlu et al.	1983	Valid.	5 !Middle Jurassic	Asia	Known from a single skull recovered from the Dashanpu Formation. Its Dorygnathus-like teeth suggest it had a piscivorous diet.
Anhanguera^[12]	Campos Kellner	1985	Valid.	7 !Early Cretaceous	S. America	A piscivorous ornithocheirid from the Santana Formation. A study of its anatomy helped resolve controversy regarding pterodacyloids' posture while on the ground.
Anurognathus^[13]	Döderlein	1923	Valid.	6 !Late Jurassic.^[14]	Europe.^[14]	A tiny (50 cm wingspan) insectivore known only from two skeletons recovered from the Solnhofen Formation.
Apatomerus^[15]	Williston	1903	Misidentification.^[2]	7 !Early Cretaceous	N. America	Probably a misidentified plesiosaur.^[2]
Arambourgiania^[16]	Nesov Kanznyshkina Cherepanov	1987	Valid.	8 !Late Cretaceous	Asia
Aralazhdarcho^[17]	Averianov	2007	Valid.	8 !Late Cretaceous	Asia
Araripedactylus	Wellnhofer	1977	Valid.	7 !Early Cretaceous	S. America
Araripesaurus	Price	1971	Valid.	7 !Early Cretaceous	S. America
Archaeoistiodactylus	Lü Fucha	2011	Valid.	5 !Middle Jurassic	Asia
Arcticodactylus	Kellner	2015	Valid.	3 !Late Triassic	Europe
Ardeadactylus^[18]	Bennett	2013	Valid.	6 !Late Jurassic	Europe
Arthurdactylus	Frey Martill	1994	Valid.	7 !Early Cretaceous	S. America
Aurorazhdarcho^[19]	Frey Meyer Tischlinger	2011	Valid.	6 !Late Jurassic.^[19]	Europe.^[19]
Aussiedraco^[20]	Kellner Rodrigues Costa	2011	Valid.	7 !Early Cretaceous	Australia
Austriadactylus^[11]	Dalla Vecchia Wild Reitner	2002	Valid.	3 !Late Triassic	Europe
Austriadraco	Kellner	2015	Valid.	3 !Late Triassic	Europe
Avgodectes^[21]	Peters	2004	Jr. synonym.	N/A	N/A	Probable junior synonym of Haopterus.
Aymberedactylus	Pêgas et al.	2016	Valid.	7 !Early Cretaceous	Asia
Azhdarcho^[22]	Nesov	1984	Valid.	8 !Late Cretaceous	Asia
Bakonydraco^[23]	Ösi Weishampel Jianu	2005	Valid.	8 !Late Cretaceous	Europe
Banguela^[24]	Headden Campos	2015	Valid.	7 !Early Cretaceous	S. America
Barbosania^[25]	Elgin Frey	2011	Valid.	8 !Late Cretaceous	S. America
Batrachognathus^[26]	Rjabinin	1948	Valid.	6 !Late Jurassic.^[27]	Asia.^[27]
Beipiaopterus^[28]	J.-C. Lü	2003	Valid.	7 !Early Cretaceous	Asia
Belonochasma^[2]	Broili	1939	Misidentification.^[2]	N/A	N/A	A non-pterosaurian gnathostome.^[2]
Bellubrunnus^[29]	Hone et al.	2012	Valid.	6 !Late Jurassic	Europe
Bennettazhia	Nesov	1991	Valid.	7 !Early Cretaceous	N. America
Bergamodactylus	Kellner	2015	Valid.	3 !Late Triassic	Europe
Bogolubovia^[30]	Nesov A. A. Yarkov	1989	Valid.	8 !Late Cretaceous	Asia
Boreopterus^[31]	Lü Qiang	2005	Valid.	7 !Early Cretaceous	Asia
Brachytrachelus	Giebel	1852	Preoccupied.	N/A	N/A	preoccupied name; now known as Scaphognathus
Brasileodactylus^[32]	Kellner	1984	Valid.	7 !Early Cretaceous	S. America
Cacibupteryx^[33]	Gasparini Fernández de la Fuente	2004	Valid.	6 !Late Jurassic	Cuba
Caiuajara^[34]	Manzig et al.	2014	Valid.	8 !Late Cretaceous	S. America
Camposipterus^[35]	Rodrigues Kellner	2013	Valid.	7 !Early Cretaceous	Europe
"Campylognathus"^[36]	Plieninger	1894	Preoccupied.	N/A	N/A	preoccupied name; now known as Campylognathoides
Campylognathoides^[11]	Strand	1928	Valid.	5 !Early Jurassic^[37]	Europe.^[37]
Carniadactylus^[38]	Dalla Vecchia	2009	Valid.	3 !Late Triassic	Europe	Formerly Eudimorphodon rosenfeldi.^[38]
Cathayopterus^[39]^[40]	Wang Zhonghe	2006	Valid.	7 !Early Cretaceous	Asia
Caulkicephalus^[41]	Steel Martill et al.	2005	Valid.	7 !Early Cretaceous	Europe
Caupedactylus^[42]	Kellner	2013	Valid.	7 !Early Cretaceous	S. America
Caviramus ^[43]	Fröbisch Fröbisch	2006	Valid.	3 !Late Triassic	Europe
Cearadactylus^[44]	Leonardi Borgomanero	1985	Valid.	7 !Early Cretaceous	S. America
Changchengopterus^[45]	Lü	2009	Valid.	6 !Late Jurassic	Asia
Chaoyangopterus^[46]	Wang Zhou	2003	Valid.	7 !Early Cretaceous	Asia
Cimoliopterus^[35]	Rodrigues Kellner	2013	Valid.	8 !Late Cretaceous	Europe
Cimoliornis	Owen	1846	Nomen nudum.	N/A	N/A
Coloborhynchus^[11]	Owen	1874	Valid.	7 !Early Cretaceous	Europe N. America S. America
Comodactylus^[47]	Galton	1981	Valid.	6 !Late Jurassic	N. America
Cretornis	Fritsch	1880	Nomen dubium.	8 !Late Cretaceous	Europe
Criorhynchus	Owen	1874	Jr. synonym.	N/A	N/A	Jr. synonym of Ornithocheirus.
Ctenochasma^[48]	von Meyer	1852	Valid.	6 !Late Jurassic.^[49]	Europe.^[49]
Cuspicephalus^[50]	Martill Etches	2013	Valid.	6 !Late Jurassic	Europe
Cycnorhamphus^[11]	Seeley	1870	Valid.	6 !Late Jurassic.^[51]	Europe.^[51]
"Daitingopterus"^[52]	Maisch Matzke Ge Sun	2004	Nomen nudum.	6 !Late Jurassic	Europe
Daohugoupterus	Cheng et al.	2015	Valid.	6 !Late Jurassic	Asia
Darwinopterus^[53]	Lü Unwin et al.	2009	Valid.	5 !Middle Jurassic	Asia	Intermediate form between rhamphorhynchoids and pterodactyloids.
Dawndraco^[54]	Kellner	2010	Valid.	8 !Late Cretaceous	N. America
Dendrorhynchoides^[55]	S.-A. Ji Q. Ji Padian	1999	Valid.	6 !Late Jurassic or 7 !Early Cretaceous	Asia
"Dendrorhynchus"^[56]	S.-A. Ji Q. Ji	1998	Preoccupied.	N/A	N/A	Preoccupied name; now known as Dendrorhynchoides.
Dermodactylus	Marsh	1881	Valid.	6 !Late Jurassic	N. America
Dimorphodon^[11]	Owen	1859	Valid.	4 !Early Jurassic^[57]	Europe.^[57]
Diopecephalus	Seeley	1871	Jr. synonym.	N/A	N/A	Jr. synonym of Pterodactylus.
Domeykodactylus^[58]	Martill Frey et al.	2000	Valid.	7 !Early Cretaceous	S. America
Doratorhynchus^[59]	Seeley	1875	Nomen vanum	6 !Late Jurassic or 7 !Early Cretaceous	Europe
Dorygnathus^[11]	Wagner	1860	Valid.	5 !Early Jurassic^[60]	Europe.^[60]
Dsungaripterus^[11]	Young	1964	Valid.	6 !Late Jurassic-7 !Early Cretaceous^[61]	Africa.^[61] Asia.^[61]
Elanodactylus^[62]	Andres Ji, Q.	2008	Valid.	7 !Early Cretaceous	Asia
Eoazhdarcho^[63]	Lü Qiang	2005	Valid.	7 !Early Cretaceous	Asia
Eopteranodon^[64]	Lü, J.C. Zhang	2005	Valid.	7 !Early Cretaceous	Asia
Eosipterus^[11]	Ji Ji	1997	Valid.	7 !Early Cretaceous	Asia
Eudimorphodon^[11]	Zambelli	1973	Valid.	2 !Late Triassic^[65]	Europe.^[65]
Europejara^[66]	Vullo et al.	2012	Valid.	7 !Early Cretaceous	Europe
Eurazhdarcho^[67]	Vremir et al.	2013	Valid.	8 !Late Cretaceous	Europe
Eurolimnornis^[68]	Kessler Jurcsák	1986	Valid.	7 !Early Cretaceous	Europe	Originally described as a bird, subsequently reinterpreted as a pterosaur.^[69]
Faxinalipterus^[70]	Bonaparte, et al.	2010	Valid.	3 !Late Triassic	S. America
Feilongus^[71]	Wang, Kellner, et al.	2005	Valid.	7 !Early Cretaceous	Asia
Fenghuangopterus^[72]	Lü Fucha Chen	2010	Valid.	5 !Middle Jurassic	Asia
Forfexopterus	Jiang et al.	2016	Valid.	7 !Early Cretaceous	Asia
Gallodactylus	Fabre	1974	Jr. synonym.	N/A	N/A	Jr. synonym of Cycnorhamphus.
Geosternbergia	Miller	1978	Jr. synonym.	N/A	N/A	Jr. synonym of Pteranodon.
Gegepterus^[73]	X. Wang Kellner et al.	2007	Valid.	7 !Early Cretaceous	Asia
Germanodactylus^[11]	Yang	1964	Valid.	6 !Late Jurassic.^[74]	Europe.^[74]
Gladocephaloideus^[75]	Lü Ji Wei Liu	2011	Valid.	7 !Early Cretaceous	Asia
Gnathosaurus^[11]	von Meyer	1833	Valid.	6 !Late Jurassic^[76]	Europe^[76]
Guidraco^[77]	Wang et al.	2012	Valid.	7 !Early Cretaceous	Asia
Gwawinapterus^[78]	Arbour Philip J. Currie	2011	Misidentification.	8 !Late Cretaceous^[78]	N. America^[78]	Initially thought to be an istiodactylid pterosaur,^[78] but subsequently reinterpreted as an indeterminate saurodontid fish.^[79]
Hamipterus^[80]	Wang et al.	2014	Valid.	7 !Early Cretaceous	Asia
Haopterus^[81]	Wang X. Lü J.	2001	Valid.	7 !Early Cretaceous	Asia
Harpactognathus^[82]	Carpenter Unwin et al.	2003	Valid.	6 !Late Jurassic	N. America
Hatzegopteryx^[83]	Buffetaut Grigorescu Csiki	2002	Valid.	8 !Late Cretaceous	Europe
Herbstosaurus^[11]	Casamiquela	1974	Valid.	6 !Late Jurassic	S. America
Hongshanopterus^[84]	X. Wang Campos et al.	2008	Valid.	7 !Early Cretaceous	Asia
Huanhepterus^[85]	Dong	1982	Valid.	6 !Late Jurassic	Asia
Huaxiapterus^[86]	J. Lü C. Yuan	2005	Valid.	7 !Early Cretaceous	Asia
Ikrandraco^[87]	Wang et al.	2014	Valid.	7 !Early Cretaceous	Asia
Ingridia^[88]	Unwin Martill	2007	Jr. synonym.	N/A	N/A	Objective Jr. synonym of Tupandactylus.
Istiodactylus^[11]	Howse Milner Martill	2001	Valid.	7 !Early Cretaceous	Europe
Jeholopterus^[89]	X. Wang Z. Zhou	2002	Valid.	5 !Middle Jurassic or 7 !Early Cretaceous	Asia
Jianchangnathus^[90]	Cheng Wang Jiang Kellner	2012	Valid.	5 !Middle Jurassic	Asia
Jianchangopterus^[91]	Lü Bo	2011	Valid.	5 !Middle Jurassic	Asia
Jidapterus^[92]	Dong Sun Wu	2003	Valid.	7 !Early Cretaceous	Asia
Kepodactylus^[93]	Harris Carpenter	1996	Valid.	6 !Late Jurassic	N. America
Kunpengopterus^[94]	X. Wang Kellner et al.	2010	Valid.	6 !Middle Jurassic	Asia
Kryptodrakon^[95]	Andres Clark Xu	2014	Valid.	5 !Middle Jurassic and/or 6 !Late Jurassic	Asia
Lacusovagus^[96]	Witton	2008	Valid.	7 !Early Cretaceous	S. America
Laopteryx	Marsh	1881	Valid.	6 !Late Jurassic	N. America
Liaoningopterus^[86]	X.-L. Wang Z.-H. Zhou	2003	Valid.	7 !Early Cretaceous	Asia
Liaoxipterus^[85]	Z. Dong J. Lü	2005	Valid.	7 !Early Cretaceous	Asia
Limnornis^[97]	Kessler Jurcsák	1984	Preoccupied.	7 !Early Cretaceous	Europe	Originally described as a bird; fossils later renamed Palaeolimnornis.
Linlongopterus^[98]	Rodrigues et alii	2015	Valid.	7 !Early Cretaceous	Asia
"Lithosteornis"^[99]	Gervais	1844	Nomen nudum.	N/A	N/A	Nomen nudum.
Lonchodectes^[11]	Hooley	1914	Valid.	7 !Early Cretaceous to 8 !Late Cretaceous	Europe
Lonchodraco^[35]	Rodrigues Kellner	2013	Valid.	7 !Early Cretaceous to 8 !Late Cretaceous	Europe
Lonchognathosaurus^[52]	Maisch Matzke Ge Sun	2004	Valid.	7 !Early Cretaceous	Asia
Longchengpterus^[100]	L. Wang L. Li et al.	2006	Valid.	7 !Early Cretaceous	Asia
Ludodactylus^[101]	Frey Martill Buchy	2003	Valid.	7 !Early Cretaceous	S. America
Maaradactylus^[102]	Bantim et alii	2014	Valid	7 !Early Cretaceous	S. America
Macrotrachelus	Giebel	1852	Jr. synonym.	N/A	N/A	Jr. synonym of Pterodactylus.
Mesadactylus^[103]	Jensen Padian	1989	Valid.	6 !Late Jurassic	N. America
Microtuban^[104]	Elgin Frey	2011	Valid.	8 !Late Cretaceous	Africa
Moganopterus^[105]	Lü et al.	2012	Valid.	7 !Early Cretaceous	Asia
Montanazhdarcho^[106]	Padian Horner de Ricqlès	1993	Valid.	8 !Late Cretaceous	N. America
Muzquizopteryx^[107]	Frey Buchy et al.	2006	Valid.	8 !Late Cretaceous	N. America
Mythunga^[108]	Molnar Thulborn	2008	Valid.	7 !Early Cretaceous	Australia
Navajodactylus^[109]	Sullivan Fowler	2011	Valid.	8 !Late Cretaceous	N. America
Nemicolopterus^[110]	X. Wang Kellner et al.	2008	Valid.	7 !Early Cretaceous	Asia
Nesodactylus^[111]	Colbert	1969	Valid.	6 !Late Jurassic	Cuba
Nesodon^[112]	Jensen Ostrom	1977	Lapsus calami.	N/A	N/A	Misspelling of Nesodactylus, also preoccupied by a toxodont.
Ningchengopterus^[113]	J. Lu	2009	Valid.	7 !Early Cretaceous	Asia
Noripterus^[11]	Yang	1973	Valid.	7 !Early Cretaceous	Asia
Normannognathus^[52]	Buffetaut J.-J. Lepage G. Lepage	1998	Valid.	6 !Late Jurassic	Europe
Nurhachius	Wang Kellner et al.	2003	Valid.	7 !Early Cretaceous	Asia
Nyctodactylus^[114]	Marsh	1881	Jr. synonym.	N/A	N/A	Jr. synonym of Nyctosaurus.
Nyctosaurus^[115]	Marsh	1876	Valid.	8 !Late Cretaceous^[116]	N. America.^[116] S. America.^[116]
"Odontorhynchus"	Stolley	1936	Jr. synonym.	N/A	N/A	Jr. synonym of Rhamphorhynchus.
"Oolithorhynchus"^[2]	Whalley	2000	Nomen nudum.^[2]			Manuscript name^[2] about which almost nothing is known.
Orientognathus	Lü et al.	2015	Valid.	6 !Late Jurassic	Asia
Ornithocephalus^[117]	von Sömmering	1812	Jr. synonym.	N/A	N/A	Jr. synonym of Pterodactylus.
Ornithocheirus^[11]	Seeley	1869	Valid.	6 !Late Jurassic.^[118]	Africa.^[118] Australia.^[118] Europe.^[118] S. America.^[118]
Ornithodesmus^[2]	Seeley	1887	Misidentification.^[2]	N/A	N/A	Misidentified dromaeosaurid.^[2]
Ornithopterus^[119]	Fitzinger	1843	Jr. synonym.	N/A	N/A	Jr. synonym of Rhamphorhynchus.
Ornithostoma^[11]	Seeley	1871	Valid.	8 !Late Cretaceous	Europe
"Osteornis"^[99]	Gervais	1844	Nomen nudum.	N/A	N/A	Nomen nudum, but appeared to be a Jr. synonym of Ornithocheirus anyway.
Pachyrhamphus^[119]	Fitzinger	1843	Preoccupied.	N/A	N/A	preoccupied name; now known as Scaphognathus
Palaeocursornis^[68]	Kessler Jurcsák	1986	Valid.	7 !Early Cretaceous	Europe	Originally described under the name Limnornis as a bird; Limnornis was preoccupied, and the fossils were subsequently reinterpreted as pterosaurian.^[69]
Palaeornis	Mantell	1844	Preoccupied.	N/A	N/A	preoccupied name.
Pangupterus	Lü et al.	2016	Valid.	7 !Early Cretaceous	Asia
Paranurognathus^[120]	Peters	2005	Jr. synonym.	N/A	N/A	Jr. synonym of Anurognathus.
Parapsicephalus^[121]	von Arthaber	1919	Valid.	4 !Early Jurassic	Europe
Peteinosaurus^[11]	Wild	1978	Valid.	2 !Late Triassic^[122]	Europe.^[122]
"Phobetor"^[2]^[11]	Bakhurina	1986	Jr. synonym.	N/A	N/A	Jr. synonym of Noripterus.^[123]
Phosphatodraco^[124]	Pereda-Suberbiola Bardet et al.	2003	Valid.	8 !Late Cretaceous	Africa
Piksi^[125]	Varricchio	2002	Valid.	8 !Late Cretaceous	N. America	Originally described as a bird, subsequently reinterpreted as a pterosaur.^[69]
Plataleorhynchus^[126]	Howse Milner	1995	Valid.	6 !Late Jurassic to 7 !Early Cretaceous	Europe
Prejanopterus^[127]	Vidarte Calvo	2010	Valid.	7 !Early Cretaceous	Europe
Preondactylus^[11]	Wild	1983	Valid.	2 !Late Triassic^[128]	Europe^[128]
"Pricesaurus"^[129]	Bonaparte Sanchez	1986	Nomen nudum.	7 !Early Cretaceous	S. America
Procoelosaurus^[130]	Atanassov	2002	Nomen ex dissertatione.^[2]
Ptenodactylus^[131]	Seeley	1869	Preoccupied	N/A	N/A	Preoccupied name.
Ptenodracon^[132]	Lydekker	1888	Jr. synonym	N/A	N/A	Junior synonym of Ctenochasma.
Pteranodon^[11]	Marsh	1876	Valid.	8 !Late Cretaceous^[133]	N. America.^[133]
Ptéro-dactyle	Cuvier	1809		N/A	N/A	Renamed Pterodactylus.
Pterodactylus	Rafinesque	1815	Valid.	6 !Late Jurassic.^[134]	Africa.^[134] Europe.^[134]
Pterodaustro^[11]	Bonaparte	1970	Valid.	7 !Early Cretaceous^[135]	S. America.^[135]
Pterofiltrus^[136]	Jiang Wang	2011	Valid.	7 !Early Cretaceous	Asia
Pteromimus^[130]	Atanassov	2002	Nomen ex dissertatione.^[2]
Pteromonodactylus	Teryaev	1967	Jr. synonym.	N/A	N/A	Jr. synonym of Rhamphorhynchus.
Pterorhynchus^[137]	Czerkas Q. Ji	2002	Valid.	6 !Late Jurassic	Asia
Pterotherium^[138]	Fischer von Waldheim	1813	Jr. synonym.	N/A	N/A	Jr. synonym of Pterodactylus.
Puntanipterus^[139]	Bonaparte Sanchez	1975	Valid.	6 !Late Jurassic to 7 !Early Cretaceous	S. America
Qinglongopterus^[140]	J. Lü Unwin et al.	2012	Valid.	5 !Middle Jurassic	Asia
Quetzalcoatlus^[11]	Lawson	1975	Valid.	8 !Late Cretaceous^[141]	N. America.^[141]
Radiodactylus^[9]	Andres Myers	2013	Valid.	7 !Early Cretaceous	N. America
Raeticodactylus^[142]	Stecher	2008	Jr. synonym	N/A	N/A	Jr. synonym of Caviramus
Rhabdopelix^[2]	Cope	1870	Misidentification.^[2]	N/A	N/A	At first it was thought to be a Triassic pterosaur, but is now known to be (at least in part) a kuehneosaurid.^[2]
Rhamphinion^[143]	Padian	1984	Valid.	4 !Early Jurassic	N. America
Rhamphocephalus^[11]	Seeley	1880	Valid.	midjur	Europe
Rhamphorhynchus^[11]	von Meyer	1846	Valid.	6 !Late Jurassic.^[144]	Africa.^[144] Europe.^[144]
Santanadactylus^[145]	de Buisonjé	1980	Valid.	7 !Early Cretaceous	S. America
Scaphognathus^[11]	Wagner	1861	Valid.	6 !Late Jurassic.^[146]	Europe.^[146]
Sericipterus	Andres Clark X. Xing	2010	Valid.	6 !Late Jurassic	Asia
Shenzhoupterus^[147]	J. Lü Unwin et al.	2008	Valid.	7 !Early Cretaceous	Asia
Sinopterus^[86]	X. Wang Z. Zhou	2003	Valid.	7 !Early Cretaceous	Asia
Siroccopteryx^[148]	Mader Kellner	1999	Jr. synonym.	N/A	N/A	Jr. synonym of Coloborhynchus.
Sordes^[11]	Sharov	1971	Valid.	6 !Late Jurassic.^[149]	Asia.^[149]
Sultanuvaisia^[8]	Nesov	1981	Misidentification.	8 !Late Cretaceous	Asia.	Actually a fish.
Tapejara^[11]	Kellner	1989	Valid.	7 !Early Cretaceous	S. America
Tendaguripterus^[150]	Unwin Heinrich	1999	Valid.	6 !Late Jurassic	Africa
Thalassodromeus^[151]	Kellner Campos	2002	Valid.	7 !Early Cretaceous	S. America
"Titanopteryx"^[152]	Arambourg	1959	Preoccupied.	8 !Late Cretaceous	Asia	Preoccupied by a simuliid blackfly. It was later renamed Arambourgiania.
Tribelesodon	Bassani	1886	Misidentification.	N/A	N/A	At first it was thought to be a Triassic pterosaur but is now known to be a misinterpreted specimen of the prolacertiform Tanystropheus.
Tropeognathus	Wellnhofer	1987	Valid.	7 !Early Cretaceous	S. America	One of the largest Ornithocheirids.
Tupandactylus^[153]	Kellner Campos	2007	Valid.	7 !Early Cretaceous	S. America
Tupuxuara^[154]	Kellner Campos	1988	Valid.	8 !Late Cretaceous	N. America S. America
Uktenadactylus^[155]	Rodrigues Kellner	2008	Disputed.	7 !Early Cretaceous	N. America
Unwindia^[156]	Martill	2011	Valid.	8 !Late Cretaceous	S. America
Utahdactylus^[157]	Czerkas Mickelson	2002	Disputed.	N/A	N/A	Probably a non-pterosaurian reptile.
Vectidraco^[158]	Naish Simpson Dyke	2013	Valid.	7 !Early Cretaceous	Europe
Volgadraco^[159]	Averianov Arkhangelsky Pervushov	2008	Valid.	8 !Late Cretaceous.	Asia.
Wenupteryx^[160]	Codorniú Gasparini	2013	Valid.	6 !Late Jurassic	S. America
Wukongopterus	X. Wang Kellner et al.	2009	Valid.	5 !Middle Jurassic	Asia
"Wyomingopteryx"^[161]	Bakker	1994	Nomen nudum.			Nomen nudum.
Yixianopterus^[162]	J. Lü S. Ji et al.	2006	Valid.	7 !Early Cretaceous	Asia	Appears to be valid, but little is known^[163]
Zhejiangopterus^[164]	Z. Cai F. Wei	1994	Valid.	8 !Late Cretaceous	Asia
Zhenyuanopterus^[165]	J. Lü	2010	Valid.	7 !Early Cretaceous	Asia

Angustinaripterus

Anhanguera

Anurognathus

Aurorazhdarcho

Bakonydraco

Campylognathoides

Cearadactylus

Coloborhynchus

Ctenochasma

Dimorphodon

Dorygnathus

Dsungaripterus

Eudimorphodon

Feilongus

Germanodactylus

Haopterus

Hatzegopteryx

Istiodactylus

Jeholopterus

Liaoningopterus

Nemicolopterus

Ningchengopterus

Noripterus

Nurhachius

Nyctosaurus

Ornithocheirus

Preondactylus

Pteranodon

Pterodactylus

Pterodaustro

Quetzalcoatlus

Scaphognathus

Sordes

Tapejara

Thalassodromeus

Tupandactylus

Tupuxuara

Zhejiangopterus

Ichnogenera

Genus	Authors	Year	Status	Age	Location	Notes
Agadirichnus	Ambroggi Lapparent	1954	Disputed.	uK	Africa
Haenamichnus^[166]	Hwang Huh et al.	2002	Valid.	uK^[166]	Asia.^[166]	More than 5 times as large as Pteraichnus, these tracks were probably made by azhdarchids.^[166]
Kouphichnium	Nopcsa	1923	Misidentification.	N/A	N/A	The trackmaker was probably a limulid.
Pteraichnus	Stokes	1957	Valid.	uJ	Asia^[167] Europe N. America	Possible junior synonym of Agadirichnus. Some tracks attributed to Pteraichnus have been considered crocodilian in origin by some scientists (Padian, 1984) however, recent studies in the past decade have overturned Padian's thoughts.
Purbeckopus	Delair	1963	Nomen dubium.	lK	Europe.

Haenamichnus

Oogenera

Although pterosaur eggs are known, some with complete embryos, no oogenera have been erected to house them. The holotype of the oospecies Oolithes sphaericus was briefly considered by Harry Govier Seeley to be pterosaurian in origin, although this attribution was dismissed before the formal erection of that oogenus.

References

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↑ Brian Andres, James Clark & Xing Xu (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–1016. doi:10.1016/j.cub.2014.03.030.
↑ Witton, Mark P. (2008). "A new azhdarchoid pterosaur from the Crato Formation (Lower Cretaceous, Aptian?) of Brazil". Palaeontology. 51 (6): 1289–1300. doi:10.1111/j.1475-4983.2008.00811.x.
↑ Kessler, E.; Jurcsák, T. (1984). "Fossil bird remains in the bauxite from Cornet (Romania, Bihor County)". Travaux du Musée d'Histoire Naturelle Grigore Antipa. 25: 393–401.
↑ Rodrigues, Taissa; Jiang, Shunxing; Cheng, Xin; Wang, Xiaolin; Kellner, Alexander W.A. (2015). "A new toothed pteranodontoid (Pterosauria, Pterodactyloidea) from the Jiufotang Formation (Lower Cretaceous, Aptian) of China and comments on Liaoningopterus gui Wang and Zhou, 2003". Historical Biology. 27 (6): 782–795. doi:10.1080/08912963.2015.1033417.
1 2 Gervais, F. L. P. 1844. Remarques sur les oiseaux fossiles. Thèse, Fac. Sci., Univ. Paris. [Abstract in Bulletin of the Société Philomathique de Paris, pp. 67-70 & L'Institut 12:293-294.]
↑ Wang, L.; Li, L.; Duan, Y.; Cheng, S.L. (2006). "A new istiodactylid pterosaur from western Liaoning, China". Geological Bulletin of China. 25 (6): 737–740.
↑ Frey, E., Martill, D., and Buchy, M. (2003). A new crested ornithocheirid from the Lower Cretaceous of northeastern Brazil and the unusual death of an unusual pterosaur. In: Buffetaut, E., and Mazin, J.-M. (eds.). Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publication 217:56-63. ISBN 1-86239-143-2.
↑ Bantim, Renan A. M.; Saraiva, Antônio A. F.; Oliveira, Gustavo R.; Sayão, Juliana M. (2014). "A new toothed pterosaur (Pterodactyloidea: Anhangueridae) from the Early Cretaceous Romualdo Formation, NE Brazil". Zootaxa. 3869 (3): 201–223. doi:10.11646/zootaxa.3869.3.1.
↑ Jensen, J. A.; Padian (1989). "Small pterosaurs and dinosaurs from the Uncompahgre fauna (Brushy Basin Member, Morrison Formation: ?Tithonian), late Jurassic, western Colorado". Journal of Paleontology. 63: 363–374.
↑ Ross A. Elgin & Eberhard Frey (2011). "A new azhdarchoid pterosaur from the Cenomanian (Late Cretaceous) of Lebanon". Swiss Journal of Geosciences. in press: 21–33. doi:10.1007/s00015-011-0081-1.
↑ Lü Junchang, Pu Hanyong, Xu Li, Wu Yanhua and Wei Xuefang (2012). "Largest toothed pterosaur skull from the Early Cretaceous Yixian Formation of Western Liaoning, China, with comments on the family Boreopteridae". Acta Geologica Sinica. 86 (2): 287–293. doi:10.1111/j.1755-6724.2012.00658.x.
↑ Padian; Horner, J.R.; de Ricqlès, A.J. (1993). "A new azhdarchid pterosaur from the Two Medicine Formation (Late Cretaceous, Campanian) of Montana, identified on the basis of bone histology". Journal of Vertebrate Paleontology. 13: 52A. doi:10.1080/02724634.1993.10011533.
↑ Frey, E.; Buchy, M.-C.; Stinnesbeck, W.; González, A.G.; Stefano, A. (2006). "Muzquizopteryx coahuilensis n.g., n. sp., a nyctosaurid pterosaur with soft tissue preservation from the Coniacian (Late Cretaceous) of northeast Mexico (Coahuila)". Oryctos. 6: 19–39.
↑ Molnar, Ralph E.; Thulborn, R.A. (2008). "An incomplete pterosaur skull from the Cretaceous of north-central Queensland, Australia". Arquivos do Museu Nacional, Rio de Janeiro. 65 (4): 461–470.
↑ Robert M. Sullivan and Denver W. Fowler (2011). "Navajodactylus boerei, n. gen., n. sp., (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (upper Campanian) of New Mexico" (PDF). Fossil Record 3. New Mexico Museum of Natural History and Science, Bulletin. 53: 393–404.
↑ Wang, X.; Kellner; Zhou, Z.; Campos, D.A. (2008). "Discovery of a rare arboreal forest-dwelling flying reptile (Pterosauria, Pterodactyloidea) from China". Proceedings of the National Academy of Sciences. 106 (6): 1983–1987. Bibcode:2008PNAS..105.1983W. doi:10.1073/pnas.0707728105. PMC 2538868. PMID 18268340.
↑ Colbert, Edwin H. (1969). "A Jurassic pterosaur from Cuba" (PDF). American Museum Novitates. 2370: 1–26. Retrieved 2007-03-03.
↑ Jensen, J. A.; Ostrom, J. H. (1977). "A second Jurassic pterosaur from North America". Journal of Paleontology. 51 (4): 867–870.
↑ Lu, J. (2009). "A baby pterodactyloid pterosaur from the Yixian Formation of Ningcheng, Inner Mongolia, China". Acta Geologica Sinica. 83 (1): 1–8. doi:10.1111/j.1755-6724.2009.00001.x.
↑ Marsh, O. C. 1881. "Note on American pterodactyls". American Journal of Science (Series 3) 21:342-343.
↑ Marsh, O. C. 1876. Principal characters of American pterodactyls. American Journal of Science, Series 3, 12:479-480.
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↑ von Sömmering, S. T. 1812. "Über einen Ornithocephalus oder über das unbekannten Thier der Vorwelt, dessen Fossiles Gerippe Collini im 5. Bande der Actorum Academiae Theodoro-Palatinae nebst einer Abbildung in natürlicher Grösse im Jahre 1784 beschrieb, und welches Gerippe sich gegenwärtig in der Naturalien-Sammlung der königlichen Akademie der Wissenschaften zu München befindet". Denkschriften der königlichen bayerischen Akademie der Wissenschaften München, mathematisch-physikalische Classe 3:89-158.
1 2 3 4 5 "Ornithocheirus." In: Cranfield, Ingrid (ed.). The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures. London: Salamander Books, Ltd. Pp. 326-329.
1 2 Fitzinger, L. J. 1843. Systema Reptilium. Braumüller et Seidel, Wien. 106 pages. [Pachyrhamphus reference on p. 35]
↑ Peters, D. 2005. Pterosaur ptrouble?. Prehistoric Times 72: 44–47.
↑ von Arthaber, G. (1919). "Studien über Flugsaurier auf Grund der Bearbeitung des Wiener exemplars von Dorygnathus banthensis Theod Sp". Denkschriften der königlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftlichen Klasse 97:391-464. [German]
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↑ "New material of dsungaripterid pterosaurs (Pterosauria: Pterodactyloidea) from western Mongolia and its palaeoecological implications." Geological Magazine, 146(5): 690-700.
↑ Pereda-Suberbiola, Xabier; Bardet, N., Jouve, S., Iarochène, M., Bouya, B., and Amaghzaz, M. (2003). "A new azhdarchid pterosaur from the Late Cretaceous phosphates of Morocco", in Buffetaut, E., and Mazin, J.-M. (eds.): "Evolution and Palaeobiology of Pterosaurs", Geological Society of London, Special Publications, 217. London: Geological Society of London, 80-90.
↑ Varricchio, David J (2002). "A new bird from the Upper Cretaceous Two Medicine Formation of Montana". Canadian Journal of Earth Sciences. 39 (1): 19–26. doi:10.1139/e01-057.
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↑ Martins Neto, R.G. (1986). Pricesaurus megalodon nov. gen. nov. sp. (Pterosauria, Pterodactyloidea), Cretaceo Inferior, chapada do Araripe (NE-Brasil). Ciência e Cultura 38(7):756-757. [Portuguese]
1 2 Atanassov, M. N. 2002. Two new archosaur reptiles from the Late Triassic of Texas. PhD dissertation, Texas Tech University, Lubbock, Texas. 352 pages.
↑ Seeley, H. G. 1869. Index to the Fossil Remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata, Arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge. 143 pages./Gray, J. E. 1845. (This book can be freely downloaded from Google Books.)
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↑ Jiang Shunxing & Wang Xiaolin (2011). "A new ctenochasmatid pterosaur from the Lower Cretaceous, western Liaoning, China". Anais da Academia Brasileira de Ciencias. 83 (4): 1243–1249. doi:10.1590/s0001-37652011000400011. ISSN 0001-3765.
↑ Czerkas, S.A., and Ji, Q. (2002). A new rhamphorhynchoid with a headcrest and complex integumentary structures. In: Czerkas, S.J. (Ed.). Feathered Dinosaurs and the Origin of Flight. The Dinosaur Museum:Blanding, Utah, 15-41. ISBN 1-932075-01-1.
↑ Fischer von Waldheim, J. G. 1813. Zoognosia tabulis synopticus illustrata, in usum praelectionum Academiae Imperialis Medico-Chirurgicae Mosquenis edita. 3rd edition, volume 1. 466 pages.
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↑ Lü, J., Unwin, D.M., Zhao, B., Gao, C. and Shen, C. (2012). "A new rhamphorhynchid (Pterosauria: Rhamphorhynchidae) from the Middle/Upper Jurassic of Qinglong, Hebei Province, China" (PDF). Zootaxa. 3158: 1–19.
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↑ Stecher, Rico (2008). "A new Triassic pterosaur from Switzerland (Central Austroalpine, Grisons), Raeticodactylus filisurensis gen. et sp. nov.". Swiss Journal of Geosciences. 101 (1): 185–201. doi:10.1007/s00015-008-1252-6. Online First
↑ Padian, K. (1984). Pterosaur remains from the Kayenta Formation (?early Jurassic) of Arizona. Palaeontology 27(2):407-413. [if you get an I/O error message, push "OK" and it should work]
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↑ Mader, B. J. and A. W. A. Kellner. 1999. "A new Anhanguerid pterosaur from the Cretaceous of Morocco". Boletim do Museu Nacional, Nova Série, Geologia 45:1-11.
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↑ Unwin, David M.; Heinrich, Wolf-Dieter (1999). "On a pterosaur jaw from the Upper Jurassic of Tendaguru (Tanzania)". Mitteilungen aus dem Museum Für Naturkunde in Berlin Geowissenschaftliche Reihe. 2: 121–134.
↑ Kellner; Campos (2002). "The function of the cranial crest and jaws of a unique pterosaur from the early Cretaceous of Brazil". Science. 297 (5580): 389–392. Bibcode:2002Sci...297..389K. doi:10.1126/science.1073186.
↑ Arambourg, C. (1959). "Titanopteryx philadelphiae nov. gen., nov. sp. Ptérosaurien géant". Notes Mém. Moyen-Orient. 7: 229–234.
↑ Campos, D.A. (2007). "Short note on the ingroup relationships of the Tapejaridae (Pterosauria, Pterodactyloidea)". Boletim do Museu Nacional. 75: 1–14.
↑ Kellner, A.W.A., and Campos, D.A. (1988). "Sobre un novo pterossauro com crista sagital da Bacia do Araripe, Cretaceo Inferior do Nordeste do Brasil. (Pterosauria, Tupuxuara, Cretaceo, Brasil)." Anais da Academia Brasileira de Ciências, 60: 459–469. [in Portuguese]
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↑ David M. Martill (2011). "A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil". Cretaceous Research. In. 32 (2): 236–243. doi:10.1016/j.cretres.2010.12.008.
↑ Czerkas, Stephen A.; Mickelson, Debra L. (2002). "The first occurrence of skeletal pterosaur remains in Utah". In Czerkas, Sylvia J. (ed.). Feathered Dinosaurs and the Origin of Flight. Blanding, Utah: The Dinosaur Museum. pp. 3–13. ISBN 1-932075-01-1.
↑ Darren Naish, Martin Simpson, Gareth Dyke (2013). "A New Small-Bodied Azhdarchoid Pterosaur from the Lower Cretaceous of England and Its Implications for Pterosaur Anatomy, Diversity and Phylogeny". PLoS ONE. 8 (3): e58451. doi:10.1371/journal.pone.0058451.
↑ Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal. 42 (6): 634–642. doi:10.1134/S0031030108060099.
↑ Laura Codorniú and Zulma Gasparini (2013). "The Late Jurassic pterosaurs from northern Patagonia, Argentina". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 399–408. doi:10.1017/S1755691013000388.
↑ Bakker, R.T. (1994) Unearthing the Jurassic. In: Science Year 1995. World Book Inc.:Chicago, London, Sydney, Toronto, 76-89. ISBN 0-7166-0595-3.
↑ Lü, J., Ji, S., Yuan, C., Gao, Y., Sun, Z. and Ji, Q. 2006. New pterodactyloid pterosaur from the Lowe Cretaceous Yixian Formation of Western Liaoning, p. 195–203. In: Lü, J., Kobayashi, Y., Huang, D. and Lee, Y. (eds), Papers form the 2005 Heyuan International Dinosaur Symposium. Beijing: Geological Publishing House.
↑ Almost new papers at the Dinosaur Mailing List describes the document in which it was named; see this page for a photograph of the type specimen
↑ Cai, Z.; Wei, F. (1994). "On a new pterosaur (Zhejiangopterus linhaiensis gen. et sp. nov.) from Upper Cretaceous in Linhai, Zhejiang, China". Vertebrata PalAsiatica. 32: 181–194.
↑ Lü, J. (2010). "A new boreopterid pterodactyloid pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, northeastern Chian". Acta Geologica Sinica. 84 (2): 241–246. doi:10.1111/j.1755-6724.2010.00204.x.
1 2 3 4 Hwang, K.-G.; et al. (2002). "'New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea'". Geological Magazine. 139 (4): 421–435. doi:10.1017/s0016756802006647.
↑ Kim, J. Y.; et al. (2006). "'The oldest record of webbed bird and pterosaur tracks from South Korea (Cretaceous Haman Formation, Changseon and Sinsu Islands): More evidence of high avian diversity in East Asia'". Cretaceous Research. 27 (1): 56–69. doi:10.1016/j.cretres.2005.10.005.

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