1991 in paleontology

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Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.^[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1991.

Arthropods

Newly named insects

Mollusca

Newly named bivalves

Archosauromorphs

• Sankar Chatterjee's discovery of a possible Triassic bird, Protoavis, if genuine, would push avian origins back almost 70 million years. The find ignites controversy over the connection between dinosaurs and birds.

Newly named dinosaurs

Data courtesy of George Olshevsky's dinosaur genera list.^[4]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Alvarezsaurus[5]	Valid taxon	Jose Bonaparte	Late Cretaceous (Santonian)	Bajo de la Carpa Formation	Argentina	An alvarezsaurid.	Amargasaurus Amurosaurus "Seismosaurus" Valdoraptor
Amargasaurus[6]	Valid taxon	Salgado and Jose Bonaparte	Early Cretaceous (Barremian)	La Amarga Formation	Argentina	An dicraeosaurid.
Amurosaurus[7]	Valid taxon	Bolotsky & Kurzanov	Late Cretaceous (Maastrichtian)	Udurchukan Formation	Argentina	A lambeosaurine hadrosaurid.
Andesaurus[8]	Valid taxon	Calvo and Bonaparte	Late Cretaceous (Santonian)	Bajo de la Carpa Formation	Argentina	An alvarezsaurid.
Becklespinax[9]	Junior synonym	George Olshevsky.	Early Cretaceous (Valanginian)	Wadhurst Clay Formation	UK	A junior objective synonym of Altispinax; new genus for "Acrocanthosaurus" altispinax Paul (1988).
Euronychodon[10]	Valid taxon	Telles-Antunes & Sigogneau-Russell	Late Cretaceous (Maastrichtian)		Portugal	A ?troodontid.
Janenschia[11]	Valid taxon	Wild	Late Jurassic (Kimmeridgian-Tithonian)	Tendaguru Formation	Tanzania	A non-titanosauriform macronarian; new genus for "Gigantosaurus" robustus E. Fraas (1908).
Protognathosaurus[9]	Valid taxon	George Olshevsky	Middle Jurassic (Bajocian)	Xiashaximiao Formation	China	A sauropod of uncertain affinities.
"Rioarribasaurus"[12]	Junior synonym	Hunt & S.G. Lucas	Late Triassic (late Norian-Rhaetian)	Chinle Formation	US	A junior synonym of Coelophysis.
"Seismosaurus"[13]	Junior synonym	Gillette
Tarascosaurus[14]	Valid taxon	Le Loeuff and Buffetaut
Taveirosaurus[10]	Valid taxon	Telles-Antunes & Sigogneau-Russell	Late Cretaceous		Portugal	An ornithischian of unknown affinities.
Tochisaurus[15]	Valid taxon	Kurzanov and Osmolska
"Ultrasauros"[9]	Junior synonym	George Olshevsky	Late Jurassic (Kimmeridgian)	Morrison Formation	US	Junior synonym of Supersaurus; replacement name for Ultrasaurus Jensen 1985 (preoccupied).
Valdoraptor[9]	Valid taxon	George Olshevsky	Early Cretaceous (late Valanginian)	Tunbridge Wells Formation	UK	An ornithomimosaur; new genus for "Megalosaurus" oweni Lydekker (1889).
Velocisaurus[5]	Valid taxon	Jose Bonaparte

Newly named birds

Genera no longer considered to be birds

• Protoavis. The avian status of Protoavis has since been almost universally rejected by paleontologists.

Pterosaurs

New taxa

Synapsids

Non-mammalian

Name	Status	Authors	Age	Unit	Location	Notes	Images
Apsisaurus[30]	Valid	Laurin	Lower Permian	Archer City Formation	USA	Formerly assigned as an eosuchian diapsid, was restudied an classified as a varanopid synapsid.[31]	Ctenorhachis
Ctenorhachis[32]	Valid	Hook Hotton	Upper Permian		USA	A sphenacodontid synapsid.

References

1. ↑ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716. 
2. ↑ MacKay, W. P. (1991). "Anochetus brevidentatus, new species, a second fossil Odontomachiti ant (Hymenoptera: Formicidae)". Journal of the New York Entomological Society. 99: 138–140. 
3. ↑ Hints, L.; Oraspõld, A.; Kaljo, D. (2000). "Stratotype of the Porkuni Stage with comments on the Röa Member (Uppermost Ordovician, Estonia)". Proceedings of the Estonian Academy of Sciences, Geology. 49 (3): 177–199. 
4. ↑ Olshevsky, George. "Dinogeorge's Dinosaur Genera List". Retrieved 2008-08-07. 
5. 1 2 Bonaparte, J.F. 1991. Los vertebrados fosiles de la Formacion Rio Colorado de Neuquen y cercanias, Cretacico Superior, Argentina. Revista del Museo Argentino de Ciencias Naturales "Bernardino Revidadia" Paleontologia 4 (3): pp. 17-23.
6. ↑ Salgado, L. and J.F. Bonaparte. 1991. Un Nuevo sauropodo Dicraeosauridae, Amargasaurus gen. et sp. nov., de la Formacion La Amarga, Neocomiano de la Provincia del Nuequen, Argentina. Ameghiniana 28: pp. 333-346.
7. ↑ Bolotsky, Yu. L. and S.K. Kurzanov. 1991. The hadrosaurs of the Amur region. In: Geology of the Pacific Border: pp.94-103.
8. ↑ Calvo, J.O. and J.F. Bonaparte, J.F. 1991. Andesaurus delgadoi gen. et sp. nov. (Saurischia-Sauropoda), dinosaurio Titanosauridae de la Formación Río Limay (Albiano-Cenomaniano), Neuquén, Argentina. Ameghiniana 28: pp. 303-310.
9. 1 2 3 4 Olshevsky, G. 1991. A Revision of the Parainfraclass Archosauria Cope, 1869, Excluding the Advanced Crocodylia. Mesozoic Meanderings 2 (1st printing): iv + 196 pages.
10. 1 2 Telles-Antunes, M. and D. Sigogneau-Russell, 1991. Nouvelles donnees sur les Dinosaures du Cretae superieur du Portugal. Compte rendu hebdomadaire des seances de l’Academie des Sciences Paris, tomo 313, seire 2: pp. 113-119.
11. ↑ Wild, R. 1991. Janenschia n. g. robusta (E.Frass 1908) pro Tornieria robusta (E. Fraas 1908) (Reptilia, Saurischia, Sauropodomorpha). Stuttgarteer Beitrage zur Naturkude Serie B (Geologie und Paleontologie) 173: pp. 1-4.
12. ↑ Hunt, A.P. and S.G. Lucas. 1991. Rioarribasaurus, a new name for a Late Triassic dinosaur from New Mexico (USA). Palaontologische Zeitschrift 6 (3/4): pp. 191-198.
13. ↑ Gillette. D.D. 1991. Seismosaurus halli, gen. et sp. nov., a new sauropod dinosaur from the Morrison Formation (Upper Jurassic/Lower Cretaceous) of New Mexico, USA. J. Vert. Paleontol. 11 (4): pp. 417-433.
14. ↑ Le Loeuff, J. and E. Buffetaut. 1991. Tarascosaurus salluvicus, new genus new species, a theropod dinosaur from the Upper Cretaceous of southern France. Geobios 24 (5): pp. 585-594.
15. ↑ Kurzanov, S.M. and H. Osmolska. 1991. Tochisaurus nemegtensis gen. et sp. n., a new troodontid (Dinosauria, Theropoda) from Mongolia. Acta Palaeont. Polonica. 36 (1): pp. 69-76.
16. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Helen F. James; Storrs L. Olson (1991). "Descriptions of Thirty-Two New Species of Birds From the Hawaiian Islands: Part II. Passeriformes" (PDF). Ornithological Monographs. 46: 1–88. doi:10.2307/40166713. 
17. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Storrs L. Olson; Helen F. James (1991). "Descriptions of Thirty-Two New Species of Birds From the Hawaiian Islands: Part I. Non-Passeriformes" (PDF). Ornithological Monographs. 45: 1–88. doi:10.2307/40166794. 
18. ↑ Lev A. Nessov; B. V. Prizemlin (1991). "Krupnyye evolyutsionno prodvinutyye neletayushchiye morskiye ptitsy odryada gesperornisoobraznykh pozdnego senona Turgayskogo proliva: pervyye nakhodki gruppy v SSSR [Large, Evolutionarily Advanced, Flightless Marine Birds of the Order Hesperornithiformes from the Late Senonian of the Turgai Strait: The first Finds of this Group in the USSR.] [in Russian, with English summ.]". Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR. 239: 85–107. 
19. ↑ Philip R. Millener; Trevor H. Worthy (1991). "Contribution to New Zealand's Late Quaternary Avifauna. II: Dendroscansor decurvirostris, a New Genus and Species of Wren (Aves: Acantisittidae)". Journal of the Royal Society of New Zealand. 21 (2): 179–200. doi:10.1080/03036758.1991.10431406. 
20. ↑ Storrs L. Olson; Helen R. James (1995). "Nomenclature of the Hawaiian Akialoas and Nukupuus (Aves: Drepanidini)" (PDF). Proceedings of the Biological Society of Washington. 108 (3): 373–387. 
21. ↑ Cécile Mourer-Chauviré (1991). "Les Horusornithidae Nov. Fam., Accipitriformes (Aves) A Articulation Intertarsienne Hyperflexible de l'Éocène du Quercy". Géobios, Mémoire spécial. 24 (supplement 1): 183–192. doi:10.1016/s0016-6995(66)80023-2. 
22. 1 2 3 4 Dénes Jánossy (1991). "Late Miocene Bird Remains from Polgárdi (W-Hungary)" (PDF). Aquila. 98: 13–35. 
23. ↑ Nikita V. Zelenkov; Andrey V. Panteleyev (2014). "Three bird taxa (Aves: Anatidae, Phasianidae, Scolopacidae) from the Late Miocene of the Sea of Azov (Southwestern Russia)". Paläontologische Zeitschrift. in press. doi:10.1007/s12542-014-0238-0. 
24. ↑ Nikita V. Zelenkov (2016). "Revision of non-passeriform birds from Polgárdi (Hungary, Late Miocene): 2. Galliformes". Paleontological Journal. 50 (6): 623–634. doi:10.1134/S0031030116060162. 
25. ↑ Thomas Schlüter (1991). "Systematik, Palökologie und Biostratonomie von Phalacrocorax kuehneanus nov. spec., einem Fossilen Kormoran (Aves: Phalacrocoracidae) aus Mutmasslich Oberpliozänen Phosporiten N-Tansanias". Berliner Geowissenschaftliche Abhandlungen, Reihe A: Geologie und Paläontologie. 134: 279–309. 
26. ↑ Jacques Cheneval; Léonard Ginsburg; Cécile Mourer-Chauviré; Benjavun Ratanasthien (1991). "The Miocene Avifauna of the Li Mae Long Locality, Thailand: Systematics and Palaeoecology". Journal of Southeast Asian Earth Sciences. 6 (2): 117–126. doi:10.1016/0743-9547(91)90103-5. 
27. ↑ Sankar Chatterjee (1991). "Cranial Anatomy and Relationships of A New Triassic Bird from Texas". Philosophical Transactions of the Royal Society of London, Series B. 332 (1265): 277–342. doi:10.1098/rstb.1991.0056. 
28. ↑ Aleksandr O. Averianov; Andrei V. Panteleyev; Olga R. Potapova; Lev A. Nessov (1991). "Lozhnozubyye ptitsy (Aves: Pelecaniformes: Odontopterygia) pozdnego paleotsena i eotsena zapadnoy okrainy drevney Azii. [Bony-toothed Birds (Aves: Pelecaniformes: Odontopterygia) of the Late Paleocene and Eocene of the Western Margin of Ancient Asia.] [in Russian, with English summ.]". Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR. 239: 3–12. 
29. ↑ Jorge I. Noriega (1991). "Un Nuevo Género de Furnariidae (Aves: Passeriformes) del Pleistoceno Inferior-Medio de la Provincia de Buenos Aires, Argentina". Ameghiniana. 28: 317–323. 
30. ↑ Laurin, M. (1991). "The osteology of a Lower permian eosuchian from Texas and a review of diapsid phylogeny". Zoological Journal of the Linnean Society. 101: 59–95. doi:10.1111/j.1096-3642.1991.tb00886.x. 
31. ↑ Reisz, R.R.; Laurin, M.; Marjanovic, D. (2010). "Apsisaurus witteri from the Lower Permian of Texas: yet another small varanopid synapsis, not a diapsid". Journal of Vertebrate Paleontology. 30 (5): 1628–1631. doi:10.1080/02724634.2010.501441. 
32. ↑ Hook, R.W.; Hotton, N. (1991). "A New Sphenacodontid Pelycosaur (Synapsida) from the Wichita Group, Lower Permian of North-Central Texas". Journal of Vertebrate Paleontology. 11 (1): 37–44. doi:10.1080/02724634.1991.10011374.