2011 in archosaur paleontology

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The year 2011 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur palaeontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2011 in paleontology included various significant developments regarding archosaurs.

This article records new taxa of fossil archosaurs of every kind that have been described during the year 2011, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2011.

Newly named crurotarsans

Name Novelty Status Authors Age Unit Location Notes Images

Acherontisuchus[1]

Gen. et sp.

Valid

  • Hastings
  • Bloch
  • Jaramillo

Middle/Late Paleocene

Cerrejón Formation

A dyrosaurid.

Anteophthalmosuchus[2]

Gen. et sp.

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A goniopholidid.

Arenysuchus[3]

Gen. et sp.

Valid

  • Puértolas
  • Canudo
  • Cruzado-Caballero

Maastrichtian

Tremp Formation

A stem crocodyloid.

Caipirasuchus[4]

Gen. et sp.

Valid

  • Iori
  • Carvalho

Turonian/Santonian

Adamantina Formation

A sphagesaurid.

Campinasuchus[5]

Gen. et sp.

Valid

Turonian/Santonian

Adamantina Formation

A baurusuchid.

Caryonosuchus[6]

Gen. et sp.

Valid

Campanian/Maastrichtian

Adamantina Formation

A sphagesaurid.

Decuriasuchus[7]

Gen. et sp.

Valid

  • França
  • Ferigolo
  • Langer

Ladinian

Santa Maria Formation

Skeletons indicate possible group behavior.

Fruitachampsa[8]

Gen. et sp.

Valid

  • Clark

Late Jurassic

Morrison Formation

A shartegosuchid.

Goniopholis kiplingi[9]

Species

Valid

Berriasian

Purbeck Limestone Group

A goniopholidid.

Goniopholis willetti[2]

Species

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A goniopholidid. Originally described as a species of Goniopholis; it was subsequently made the type species of a separate genus Hulkepholis.[10]

Labidiosuchus[11]

Gen. et sp.

Valid

Maastrichtian

Marília Formation

A notosuchian.

Leiokarinosuchus[2]

Gen. et sp.

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A neosuchian.

Lorosuchus[12]

Gen. et sp.

Valid

  • Pol
  • Powell

Paleocene

Rio Loro Formation

A sebecid.

Neptunidraco[13]

Gen. et sp.

Valid

  • Cau
  • Fanti

late Bajocian/early Bathonian

Rosso Ammonitico Veronese Formation

The oldest known metriorhynchid.

Pepesuchus[14]

Gen. et sp.

Valid

Campanian/Maastrichtian

Presidente Prudente Formation

A peirosaurid.

Pietraroiasuchus[15]

Gen. et sp.

Valid

Early Albian

Pietraroia Plattenkalk

A hylaeochampsid.

Pissarrachampsa[16]

Gen. et sp.

Valid

  • Montefeltro
  • Larsson
  • Langer

Campanian/Maastrichtian

Vale do Rio do Peixe Formation

A baurusuchid. The type species is Pissarrachampsa sera.

Theriosuchus grandinaris[17]

Species

Valid

  • Lauprasert
  • Laojumpon
  • et al.[CAL 7]

Berriasian/Barremian

Khua Formation

A fifth species of Theriosuchus.

Newly named non-avian dinosaurs

Name Novelty Status Authors Age Unit Location width="33%" Notes Images

Acristavus[18]

Gen. et sp.

Valid

Campanian

Two Medicine Formation
Wahweap Formation

A brachylophosaurin hadrosaurine.

Ahshislepelta[19]

Gen. et sp.

Valid

  • Burns
  • Sullivan

Late Campanian

Kirtland Formation

An ankylosaurine ankylosaurid.

Albinykus[20]

Gen. et sp.

Valid

Late Cretaceous

An alvarezsaurid theropod.

Amtocephale[21]

Gen. et sp.

Valid

  • Watabe
  • Tsogtbaatar
  • Sullivan

Turonian-Santonian

Baynshire Formation

A pachycephalosaurid.

Angolatitan[22]

Gen. et sp.

Valid

Late Turonian

Itombe Formation

A somphospondylian sauropod.

Arcusaurus[23]

Gen. et sp.

Valid

  • Yates
  • Bonnan
  • Neveling

Lower Jurassic

Elliot Formation

Basal Sauropodomorph

Atacamatitan[24]

Gen. et sp.

Valid

Late Cretaceous

Tolar Formation

A titanosaur.

Brontomerus[25]

Gen. et sp.

Valid

  • Taylor
  • Wedel
  • Cifelli

Aptian/Albian

Cedar Mountain Formation

A camarasauromorph sauropod.

Daemonosaurus[26]

Gen. et sp.

Valid

Late Triassic

Chinle Formation

A primitive theropod.

Delapparentia[27]

Gen. et sp.

Valid

  • Ruiz-Omeñaca

Early Barremian

Camarillas Formation

An iguanodontoid.

Demandasaurus[28]

Gen. et sp.

Valid

  • Fernández-Baldor
  • Canudo
  • et al.[CAL 13]

Late Barremian-early Aptian

Castrillo la Reina Formation

A rebbachisaurid sauropod.

Drusilasaura[29]

Gen. et sp.

Valid

  • Navarrete
  • Casal
  • Martínez

Cenomanian/Turonian

Bajo Barreal Formation

A possible lognkosaurian titanosaur.

Eodromaeus[30]

Gen. et sp.

Valid

Middle Carnian

Ischigualasto Formation

A basal theropod.

Epichirostenotes[31]

Gen. et sp.

Valid

  • Sullivan
  • Jasinski
  • Van Tomme

Late Campanian

Horseshoe Canyon Formation

A caenagnathid.

Haya[32]

Gen. et sp.

Valid

Late Cretaceous

Javkhlant Formation

Basal ornithopod with many gastroliths.

Jaklapallisaurus[33]

Gen. et. sp.

Valid

Late Norian-earliest Rhaetian

Upper Maleri Formation
Lower Dharmaram Formation

A plateosaurid.

Koreaceratops[34]

Gen. et sp.

Valid

  • Lee
  • Ryan
  • Kobayashi

Albian

"Tando beds"

A basal ceratopsian.

Koreanosaurus[35]

Gen. et sp.

Valid

?Santonian/Campanian

Seonso Conglomerate

A hypsilophodontid ornithopod.

Leonerasaurus[36]

Gen. et sp.

Valid

  • Pol
  • Garrido
  • Cerda

Pliensbachian/Toarcian

Las Leoneras Formation

An anchisaurian sauropodomorph.

Leyesaurus[37]

Gen. et sp.

Valid

Lower Jurassic

Quebrada del Barro Formation

A massospondylid sauropodomorph.

Linhenykus[38]

Gen. et sp.

Valid

Campanian

Wulansuhai Formation

An alvarezsaurid.

Linhevenator[39]

Gen. et sp.

Valid

Campanian

Bayan Mandahu Formation

An short-armed troodontid.

Manidens[40]

Gen. et sp.

Vald

  • Pol
  • Rauhut
  • Becerra

Aalenian/Early Bathonian

Cañadón Asfalto Formation

A heterodontosaurid.

Nambalia[33]

Gen. et. sp.

Valid

Late Norian-earliest Rhaetian

Upper Maleri Formation

A basal sauropodomorph.

Narambuenatitan[41]

Gen. et sp.

Valid

  • Filippi
  • García
  • Garrido

lower-middle Campanian

Anacleto Formation

A lithostrotian titanosaur.

Ojoraptorsaurus[31]

Gen. et sp.

Valid

  • Sullivan
  • Jasinski
  • Van Tomme

Lower Maastrichtian

Ojo Alamo Formation

A caenagnathid.

Omeisaurus jiaoi[42]

Species

Valid

Middle Jurassic

Xiashaximiao Formation

A species of Omeisaurus.

Osmakasaurus[43]

Gen. et comb.

Valid

  • McDonald

Valanginian

Lakota Formation

A new genus for "Camptosaurus" depressus (Gilmore, 1909).

Oxalaia[44]

Gen. et sp.

Valid

Cenomanian

Alcântara Formation

Huge spinosaur 12–14 m long.

Pampadromaeus[45]

Gen. et sp.

Valid

Carnian

Santa Maria Formation

A very basal sauropodomorph.

Pamparaptor[46]

Gen. et sp.

Valid

  • Porfiri
  • Calvo
  • Santos

Turonian/Coniacian

Portezuelo Formation

A deinonychosaur.

Petrobrasaurus[47]

Gen. et sp.

Valid

Santonian

Plottier Formation

A titanosaur.

Propanoplosaurus[48]

Gen. et sp.

Valid

  • Stanford
  • Weishampel
  • Deleon

Early Aptian

Patuxent Formation

A nodosaurid.

Qiupalong[49]

Gen. et sp.

Valid

Late Cretaceous

Qiupa Formation

An ornithomimid.

Ratchasimasaurus[50]

Gen. et sp.

Valid

  • Shibata
  • Jintasakul
  • Azuma

Aptian

Khok Kruat Formation

A non-hadrosaurid iguanodontian.

Sarahsaurus[51]

Gen. et sp.

Valid

Sinemurian/Pliensbachian

Kayenta Formation

A basal sauropodomorph

Siamodon[52]

Gen. et sp.

Valid

  • Buffetaut
  • Suteethorn

Aptian

Khok Kruat Formation

An advanced iguanodontian.

Spinops[53]

Gen. et sp.

Valid

Campanian

Oldman or Dinosaur Park Formation

A centrosaurine ceratopsian.

Stegoceras novomexicanum[54]

Species

Valid

  • Jasinski
  • Sullivan

Late Campanian

Fruitland Formation
Kirtland Formation

Second known species of Stegoceras.

Talos[55]

Gen. et sp.

Valid

Late Campanian

Kaiparowits Formation

A troodontid theropod.

Tapuiasaurus[56]

Gen. et sp.

Valid

Aptian

Quiricó Formation

A titanosaur.

Teratophoneus[57]

Gen. et sp.

Valid

Late Campanian

Kaiparowits Formation

A tyrannosaurine tyrannosaurid.

Thescelosaurus assiniboiensis[58]

Species

Valid

  • Brown
  • Boyd
  • Russell

Late Maastrichtian

Frenchman Formation

Third known species of Thescelosaurus.

Titanoceratops[59]

Gen. et sp.

Valid

  • Longrich

Late Campanian

Fruitland or Kirtland Formation

A giant chasmosaurine ceratopsian from New Mexico.

Traukutitan[60]

Gen. et sp.

Valid

  • Juárez Valieri
  • Calvo

Santonian

Bajo de la Carpa Formation

A possible lognkosaurian titanosaur.

Uteodon[43]

Gen. et comb.

Valid

  • McDonald

Late Jurassic

Morrison Formation

A new genus for "Camptosaurus" aphanoecetes (Carpenter & Wilson, 2008).

Veterupristisaurus[61]

Gen. et sp.

Valid

  • Rauhut

Late Kimmeridgian/earliest Tithonian

Tendaguru Formation

A carcharodontosaurid.

Xiaotingia[62]

Gen. et sp.

Valid

early Late Jurassic

Tiaojishan Formation

An Archaeopteryx-like paravian.

Xuwulong[63]

Gen. et sp.

Valid

  • You
  • Li
  • Liu

Early Cretaceous

Xinminpu Group

A hadrosauriform.

Zhuchengtyrannus[64]

Gen. et sp.

Valid

Late Cretaceous

Wangshi Group

A tyrannosaurine tyrannosaurid.

Newly named birds

Name Novelty Status Authors Age Unit Location Notes Images

Alca carolinensis [65]

Sp. nov

Valid

N. Adam Smith

Julia A. Clarke

Early Pliocene

Yorktown Formation

 USA:

 North Carolina

An Alcidae related to the razorbill.

Alca minor [65]

Sp. nov

Valid

N. Adam Smith

Julia A. Clarke

Early Pliocene

Yorktown Formation

 USA:

 North Carolina

An Alcidae related to the razorbill.

Alca olsoni [65]

Sp. nov

Valid

N. Adam Smith

Julia A. Clarke

Early Pliocene

Yorktown Formation

 USA:

 North Carolina

An Alcidae related to the razorbill.

Alectroenas payandeei [66]

Sp. nov

Valid

Julian Pender Hume

Holocene-Recent

 Mauritius

A pigeon, Columbidae from Rodrigues Island.

?Aptornis proasciarostratus [67]

Sp. nov.

Valid

Trevor H. Worthy

Alan J. D. Tennyson

R. Paul Scofield

Early Miocene,

Altonian

Manuherikia Group,

St Bathans Fauna

 New Zealand

The earliest known Aptornithidae Bonaparte, 1856.

Ardea sytchevskayae [68]

Sp. nov.

Valid

Nikita V. Zelenkov

Middle Miocene

 Mongolia

An Ardeidae, earliest member of Ardea

Aurorornis taurica[69]

Gen. et sp. nov

Valid

Andrey V. Panteleyev

Eocene (Lutetian)

Crimean Peninsula

An owl belonging to the family Protostrigidae. The type species of the new genus.

Australlus gagensis [70]

Sp. nov.

Valid

Trevor H. Worthy

Walter E. Boles

Oligocene-Miocene

Riverleigh

 Australia:

 Queensland

A Rallidae. The type species is Gallinula disneyi Boles, 2005.

Bohaiornis guoi [71]

Gen. nov. et Sp. nov.

Valid

Hu Dongyu

Li Li

Hou Liannaim

Xu Xing

Barremian/Aptian

Yixian Formation

 China

An Enantiornithes Walker, 1981, Bohaiornithidae Wang, Zhou, O’Connor et Zelenkov, 2014.

Buteo praebuteo [72]

Sp. nov

Valid

Denis Sobolev in Sobolev & Marisova

Miocene

 Ukraine

An Accipitridae, a species of Buteo.

Buteo sarmathicus [72]

Sp. nov

Valid

Denis Sobolev in Sobolev & Marisova

Miocene

 Ukraine

An Accipitridae, a species of Buteo.

Columba thiriouxi [66]

Sp. nov

Valid

Julian Pender Hume

Holocene-Recent

 Mauritius

A pigeon, Columbidae from Mauritius.

Colymbiculus udovinchenkoi [73]

Gen. et sp. nov

Valid

Gerald Mayr

Evgenij Zvonok

Middle Eocene

Early Lutetian

 Ukraine

A Gaviidae, this is the type species of the new genus.

Cyrilavis colburnorum [74]

Species

Valid

Daniel T. Ksepka

Julia A. Clarke

Lance Grande

Eocene

Green River Formation

 USA:

 Wyoming

A Halcyornithidae Harrison et Walker, 1972, a stem Psittaciformes.

Eostrix tsaganica [75]

Sp. nov

Valid

Evgeney N. Kurochkin

Gareth J. Dyke

Early Eocene

 Mongolia

A Protostrigidae Wetmore, 1933.

Falco bulgaricus [76]

Sp. nov

Valid

Zlatozar N. Boev

Late Miocene

 Bulgaria

A Falconidae.

Feducciavis loftini [77]

Gen. nov. et Sp. nov.

Valid

Storrs L. Olson

Middle Miocene

Calvert Formation

 USA:

 Virginia

A Sternidae close to Anous and Procelsterna.

Gallirallus steadmani [78]

Sp. nov.

Valid

Trevor H. Worthy

Robert Bollt

Quaternary

Subrecent

 French Polynesia

A Rallidae as big as Gallirallus philippensis from Tubuai, Austral Islands.

Gracilornis jiufotangensis [79]

Gen. et sp.

Valid

Li Li

Hou Shi-Lin

Aptian

Jiufotang Formation

 China

An Enatiornithes Walker, 1981.

Heterostrix tatsinensis [75]

Gen. et sp. nov

Valid

Evgeney N. Kurochkin

Gareth J. Dyke

Early Oligocene

Hsanda Gol Formation

 Mongolia

A Heterostrigidae Fam. nov. Kurochkin & Dyke, 2011, this is the type species of the new genus and the type genus of the new family.

Hoazinavis lacustris [80]

Gen. et sp.

Valid

Gerald Mayr

Herculano M. F. de Alvarenga

Cécile Mourer-Chauviré

Late Oligocene/Early Miocene

Tremembé Formation

 Brazil

An early Opisthocomidae.

Itaboravis elaphrocnemoides [81]

Gen. et sp.

Valid

Gerald Mayr

Herculano M. F. de Alvarenga

Julia A. Clarke

Early late Paleocene

Itaboraian

 Brazil

A possible member of Cariamae. This is the type species of the new genus.

Jamna szybiaki [82]

Gen. et sp.

Valid

Zbigniew M. Bochenski

Teresa Tomek

Malgorzata Bujoczek

Krzysztof Wertz

Oligocene

Carpathian flysch

 Poland

An early Passeriformes.

Kaiika maxwelli [83]

Gen. et sp.

Valid

R. Ewan Fordyce

Daniel Thomas

Early Eocene

Kauru Formation

 New Zealand

A basal Spheniscidae.

Laricola intermedia [84]

Sp. nov.

Valid

Vanessa L. De Pietri

Lodc Costeur

Marcel Güntert

Gerald Mayr

Early Miocene

MN 2a

 France

A Charadriiformes, Laricolidae De Pietri, Costeur, Güntert et G. Mayr, 2011.

Laricola robusta [84]

Sp. nov.

Valid

Vanessa L. De Pietri

Lodc Costeur

Marcel Güntert

Gerald Mayr

Early Miocene

MN 2a

 France

A Charadriiformes, Laricolidae De Pietri, Costeur, Güntert et G. Mayr, 2011.

Lavocatavis africana [85]

Gen. et sp.

Valid

Cécile Mourer-Chauviré

Rodolphe Tabuce

M’hammed Mahboubi

Mohammed Adaci

Mustapha Bensalah

Eocene

Glib Zegdou Formation

 Algeria

A Phorusrhacoidea.

Luscinia jurcsaki [86]

Sp. nov

Valid

Eugen Kessler

Márton Venczel

Middle Miocene

MN 6-8

 Romania

A member of the Muscicapidae

Mancalla lucasi [87]

Sp. nov

Valid

N. Adam Smith

Pliocene or Early Pleistocene

Niguel Formation

San Diego Formation

 USA:

 California

A Charadriiformes, Alcidae, Mancallinae L. H. Miller, 1946.

Mancalla vegrandis [87]

Sp. nov

Valid

N. Adam Smith

Pliocene or Early Pleistocene

San Diego Formation
San Mateo Formation

 USA:

 California

A Charadriiformes, Alcidae, Mancallinae L. H. Miller, 1946.

Miomancalla howardae [87]

Gen. et sp. et comb. nov

Valid

N. Adam Smith

Late Miocene to Early Pliocene

Capistrano Formation
San Mateo Formation

 USA:

 California

A Charadriiformes, Alcidae, Mancallinae L. H. Miller, 1946, this is the type species of the new genus, that also contains "Praemancalla" wetmorei Howard (1966).

Mystiornis cyrili [88]

Gen. et sp.

Valid

Evgeny N. Kurochkin

Nikita V. Zelenkov

Alexandr O. Averianov

Sergei V. Leshchinskiy

Barremian/Aptian

Shestakovo-1 locality

 Russian Federation

Namaortyx sperrgebietensis [89]

Gen. nov. et Sp. nov.

Valid

Cécile Mourer-Chauviré

Martin Pickford

Brigitte Senut

Middle Eocene

Probably Early Lutetian

 Namibia

A Galliformes Incertae Sedis. This is the type species of the new genus.

Nambashag billerooensis [90]

Gen. nov. et Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene/Early Miocene

Etadunna Formation

 Australia:

 South Australia

A Phalacrocoracidae, this is the type species of the new genus.

Nambashag microglaucus [90]

Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene/Early Miocene

Namba Formation

 Australia:

 South Australia

A Phalacrocoracidae.

Nelepsittacus minimus [91]

Gen. nov. et sp. nov.

Valid

Trevor H. Worthy

Alan J. D. Tennyson

R. Paul Scofield

Early Miocene

St. Bathans Fauna,

Manuherikia Section

 New Zealand

A Psittaciformes, Nestoridae, smallest species of the genus and the type species of the new genus.

Nelepsittacus donmertoni [91]

Sp. nov.

Valid

Trevor H. Worthy

Alan J. D. Tennyson

R. Paul Scofield

Early Miocene

St. Bathans Fauna,

Manuherikia Section

 New Zealand

A Psittaciformes, Nestoridae.

Nelepsittacus daphneleeae [91]

Sp. nov.

Valid

Trevor H. Worthy

Alan J. D. Tennyson

R. Paul Scofield

Early Miocene

St. Bathans Fauna,

Manuherikia Section

 New Zealand

A Psittaciformes, Nestoridae.

Nesoenas cicur [66]

Sp. nov

Valid

Julian P. Hume

Holocene-Recent

 Mauritius

An pigeon, Columbidae from Mauritius.

Nogusunna conflictoides [92]

Gen. nov et sp. nov

Valid

Nikita V. Zelenkov

Middle Miocene

Oshin Formation

 Mongolia

A diving duck, Anatidae.

Parahongshanornis chaoyangensis [93]

Gen. et sp. nov.

Valid

Li Li

Wang Jing-Qi

Hou Shi-Lin

Early Cretaceous

Jiufotang Formation

 China

A Hongshanornithidae O’Connor, Gao et Chiappe, 2010.

Pellornis mikkelseni [94]

Gen. et sp. nov

Valid

Sara Bertelli

Luis M. Chiappe

Gerald Mayr

Earliest Eocene

Fur Formation

 Denmark

A Messelornithidae Hesse, 1988, this is the type species of the new genus.

Piculoides saulcetensis [95]

Gem. nov. et sp. nov.

Valid

Vanessa L. De Pietri

Albrecht Manegold

Lodc Costeur

Gerald Mayr

Early Miocene

MN 2

 France

A Picidae, this is the type species of the new genus.

Protomelanitta gracilis [92]

Gen. nov et sp. nov

Valid

Nikita V. Zelenkov

Middle Miocene

Oshin Formation

 Mongolia

A diving duck, Anatidae, this is the type species of the new genus, a second species of the genus is Protomelanitta shihuibas (Hou, 1985), formally known as Aythya shihuibas Hou, 1985.

Qiliania graffini [96]

Gen. et sp.

Valid

Ji Shu-An

Jessie Atterholt

Jingmai K. O’Connor

Matthew C. Lamanna

Jerald D. Harris

Li Da-Qing

You Hai-Lu

Peter Dodson

Aptian

Xiagou Formation

 China

An Enantiornithes Walker, 1981.

Sharganetta mongolica [92]

Gen. nov et sp. nov

Valid

Nikita V. Zelenkov

Middle Miocene

Oshin Formation

 Mongolia

A diving duck, Anatidae, this is the type species of the new genus.

Sternalara milneedwardsi [84]

Sp. nov.

Valid

Vanessa L. De Pietri

Lodc Costeur

Marcel Güntert

Gerald Mayr

Early Miocene

MN 2a

 France

A Charadriiformes, family Incertae Sedis.

Sternalara minuta [84]

Gen. nov. et sp. nov.

Valid

Vanessa L. De Pietri

Lodc Costeur

Marcel Güntert

Gerald Mayr

Early Miocene

MN 2a

 France

A Charadriiformes, family Incertae Sedis this is the typespecies of the genus.

Tonsala buchanani [97]

Sp. nov

Valid

Gareth J. Dyke

Xia Wang

Michael B. Habib

Early Oligocene, probably also late Eocene

Makah Formation

Pysht Formation

 USA:

 Washington

A Plotopteridae Howard, 1969, a species of Tonsala.

Newly named pterosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Aurorazhdarcho[98]

Gen. et sp.

Valid

  • Frey
  • Meyer
  • Tischlinger

Tithonian

Solnhofen limestone

An azhdarchoid[98] or a ctenochasmatid.[99] The type species, Aurorazhdarcho primordius, was subsequently found to be synonymous with "Pterodactylus" micronyx von Meyer (1856); however, Aurorazhdarcho was retained as a distinct genus, creating a new combination Aurorazhdarcho micronyx.[99]

Aussiedraco[100]

Gen. et sp.

Valid

  • Kellner
  • Rodrigues
  • Costa

Albian

Toolebuc Formation

An ornithocheiroid.

Barbosania[101]

Gen. et sp.

Valid

  • Elgin
  • Frey

Cenomanian

Santana Formation

An ornithocheirid.

Darwinopterus robustodens[102]

Species

Valid

Middle Jurassic

Tiaojishan Formation

A darwinopterid.

Jianchangopterus[103]

Gen. et sp.

Valid

  • Bo

Middle Jurassic

Tiaojishan Formation

A rhamphorhynchid.

Navajodactylus[104]

Gen. et sp.

Valid

  • Sullivan
  • Fowler

Late Campanian

Kirtland Formation
Dinosaur Park Formation

A possible azhdarchid.

Pterofiltrus[105]

Gen. et sp.

Valid

  • Jiang
  • Wang

Early Cretaceous

Yixian Formation

A ctenochasmatid.

Unwindia[106]

Gen. et sp.

Valid

  • Martill

Cenomanian

Santana Formation

A basal ctenochasmatoid.

Notes

  1. Carvalho, Teixeira, Ferraz, Ribeiro, Martinelli, Neto, Sertich, Cunha, Cunha, Ferraz.
  2. Kellner, Campos, Riff, de Andrade.
  3. de Andrade, Edmonds, Benton, Schouten.
  4. Kellner, Figueiredo, Azevedo, Campos.
  5. Campos, Oliveira, Figueiredo, Riff, Azevedo, Carvalho, Kellner
  6. Buscalioni, Piras, Vullo, Signore, Barbera
  7. Lauprasert, Laojumpon, Saenphala, Cuny, Thirakhupt, Suteethorn
  8. Gates, Horner, Hanna, Nelson.
  9. Nesbitt, Clarke, Turner, Norell.
  10. Mateus, Jacobs, Schulp, Polcyn, Tavares, Neto, Morais, Antunes.
  11. Kellner, Rubilar-Rogers, Vargas, Suárez
  12. Sues, Nesbitt, Berman, Henrici
  13. Fernández-Baldor, Canudo, Huerta, Montero, Pereda Suberbiola, Salgado.
  14. Martinez, Sereno, Alcober, Colombi, Renne, Montañez, Currie.
  15. Makovicky, Kilbourne, Sadleir, Norell.
  16. 1 2 Novas, Ezcurra, Chatterjee, Kutty.
  17. Huh, Lee, Kim, Lim, Godefroit.
  18. Apaldetti, Martinez, Alcober, Pol.
  19. Xu, Sullivan, Pittman, Choiniere, Hone, Upchurch, Tan, Xiao, Tan, Han.
  20. Xu, Tan, Sullivan, Han, Xiao.
  21. Jiang, Li, Peng, Ye
  22. Kellner, Azvedo, Machado, Carvalho, Henriques.
  23. Schultz, Bittencourt, Soares, Fortier, Silva, Langer.
  24. Filippi, Canudo, Salgado, Garrido, Garcia, Cerda, Otero.
  25. Xu, Kobayashi, Lü, Lee, Liu, Tanaka, Zhang, Jia, Zhang.
  26. Farke, Ryan, Barrett, Tanke, Braman, Loewen, Graham.
  27. Zanno, Varricchio, O'Connor, Titus, Knell.
  28. Zaher, Pol, Carvalho, Nascimento, Riccomini, Larson, Juarez-Valieri, Pires-Domingues, da Silva Jr., Campos.
  29. Carr, Williamson, Britt, Stadtman.
  30. Xu, You, Du, Han.
  31. Hone, Wang, Sullivan, Zhao, Chen, Li, Ji, Ji, Xu.
  32. Lü, Xu, Chang, Zhang.

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