Penion
| Penion Temporal range: Early Paleocene to Recent, 66.0–0.0 Ma | |
|---|---|
 | |
| A shell of Penion maximus | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| (unranked): | clade Caenogastropoda clade Hypsogastropoda |
| Superfamily: | Buccinoidea |
| Family: | Buccinidae |
| Genus: | Penion Fischer, 1884[1] |
| Species | |
|
See text. | |
| Synonyms | |
Penion is a genus of large sea marine snails, commonly known as siphon whelks, classified within the mollusc family Buccinidae, the true whelks.
Description
Siphon whelks are large, benthic marine snails, or whelks.[4] Most species inhabit soft sediments on the continental shelf.[5] Small shells and fossils of Penion can sometimes be confused with those of Aeneator.[6]
Distribution
Penion species are restricted to the Southern Hemisphere. Two extant species are currently classified in Australia[7][8] and six extant species are documented in New Zealand.[4]
Numerous fossil species are recorded in New Zealand,[6] Australia,[7] Argentina and Chile,[9][10] and Antarctica.[11][12]
Evolution
Penion is currently classified within Buccinidae, the family of true whelks.[13] A molecular phylogeny of Buccinidae based on the mitochondrial 16S rRNA gene suggested that Penion is a sister clade to Kelletia, a genus of whelks with extant species found in the Sea of Japan and in waters off of California, United States and Mexico.[13]
Species
Species and subspecies in the genus Penion include:
- † Penion affixus (Finlay, 1930)[6]
- † Penion antarctocarinatus (Stilwell & Zinsmeister, 1992)[11]
- † Penion asper (Marwick, 1928)[6]
- † Penion australocapax (Stilwell & Zinsmeister, 1992)[11][12]
- † Penion bartrumi (Laws, 1941)[6]
- Penion benthicolus (Dell, 1956)[4][6]
- † Penion brazieri (Fleming, 1955)[6]
- Penion chathamensis (Powell, 1938)[13]
- † Penion clifdenensis (Finlay, 1930)[6]
- † Penion crassus (Frassinetti, 2000)[9]
- † Penion crawfordi (Hutton, 1873)[6]
- Penion cuvierianus (Powell, 1927)[4][6]
- † Penion darwinianus (Philippi, 1887)
- † Penion diversum (Frassinetti, 2000)[9]
- † Penion domeykoanus (Philippi, 1887)
- † Penion exoptatus (Powell & Bartrum, 1929)[6]
- Penion fairfieldae (Powell, 1947)[4][6]
- † Penion finlayi (Laws, 1930)[6]
- † Penion gauli (Marwick, 1928)[6]
- † Penion haweraensis (Powell, 1931)[6]
- † Penion hiatulus (Powell, 1947)[6]
- † Penion huttoni (L.R. King, 1934)[6]
- † Penion imperfectus (Powell, 1947)[6]
- † Penion interjunctus (Finlay, 1930)[6]
- † Penion koruahinensis (Powell & Bartrum, 1928)[6]
- † Penion longirostris (Tate, 1888)[7]
- † Penion macsporrani (Philippi, 1887)
- Penion mandarinus (Duclos, P.L., 1831)[7]
- Penion mandarinus waitei (Hedley, 1903)
- † Penion marwicki (Finlay, 1930)[6]
- Penion maximus (Tryon, G.W., 1881)[7]
- † Penion oncodes (Philippi, 1887)
- Penion ormesi (Powell, 1927)
- † Penion parans (Finlay, 1930)[6]
- † Penion patagonensis (Reichler, 2010)[10]
- † Penion petitianus (d'Orbigny, 1842)
- † Penion proavitus (Finlay & Marwick, 1937)[6]
- † Penion roblini (Tenison Woods, 1876)[7]
- † Penion spatiosus (Tate, 1888)[7]
- † Penion subrectus (Ihering, 1899)
- † Penion subreflexus (G.B. Sowerby I, 1846)
- † Penion subregularis (d'Orbigny, 1852)
- Penion sulcatus (Lamarck, 1816)[4][6][13]
- † Penion winthropi (Marwick, 1965)[6]
References
- ↑ Fischer, P. 1884. Manuel de Conchyliologie et de Paléontologie Conchyliologique ou histoire naturelle des mollusques vivant et fossiles. Paris : F. Savy Vol. 7 pp. 609-688.
- ↑ Iredale, T. 1924. Results from Roy Bell's molluscan collections. Proceedings of the Linnean Society of New South Wales 49(3): 179-279, pls 33-36.
- ↑ Iredale, T. 1914. On some invalid molluscan generic names. Proceedings of the Malacological Society of London 11: 170-178.
- 1 2 3 4 5 6 7 Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
- ↑ Dell, R.K.. 1962. New Zealand Marine Provinces - do they exist? Tuatara, 10: 43 - 52. Online Copy courtesy of New Zealand Electronic Text Collection
- 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Beu, A.G. and Maxwell, P.A. 1990. Cenozoic Mollusca of New Zealand. New Zealand Geological Survey Bulletin, 58.
- 1 2 3 4 5 6 7 Ponder, W.F.. 1973. A review of the Australian species of Penion Fischer (Neogastropoda: Buccinidae). Journal of the Malacological Society of Australia 2: 401–428.
- ↑ Wilson, B. 1994. Australian Marine Shells. Prosobranch Gastropods. Kallaroo, WA : Odyssey Publishing Vol. 2 370 pp.
- 1 2 3 Frassinetti, D.C. 2000. Upper Pliocene marine molluscks from Guafo Island, southern Chile. Part II. Gastropoda. Boletin del Museo Nacional de Historia Natural, Chile 49: 131–161.
- 1 2 Reichler, V.A. 2010. Estratigrafía y paleontología del Cenozoico marino del Gran Bajo y Salinas del Gualicho, Argentina y descripción de 17 especies nuevas. Andean Geology, 37(1): 177-219
- 1 2 3 Stilwell, J.D., Zinsmeister, W.J. 1992. Molluscan systematics and biostratigraphy, lower Tertiary La Meseta Formation, Seymour Island, Antarctic Peninsula. American Geophysical Union Antarctica Research Series 55: 126-128. DOI: 10.1029/AR055 ISBN 9781118667705
- 1 2 Beu, A.G. 2009. Before the ice: Biogeography of Antarctic Paleogene molluscan faunas. Palaeogeography, Palaeoclimatology, Palaeoecology 284: 191–226. doi:10.1016/j.palaeo.2009.09.025
- 1 2 3 4 Hayashi, S. 2005. The molecular phylogeny of the Buccinidae (Caenogastropoda: Neogastropoda) as inferred from the complete mitochondrial 16s rRNA gene sequences of selected representatives. Molluscan Research, 25: 85–98.
External links
- World Register of Marine Species: Penion P. Fischer, 1884
- Museum of New Zealand Te Papa Tongarewa, Taxon: Penion (Genus)
- Fossilworks Database Penion Fischer 1884 (true whelk)
- Revised descriptions of New Zealand Cenozoic Mollusca from Beu and Maxwell (1990)
- Checklist of the Recent Mollusca Recorded from the New Zealand Exclusive Economic Zone
Further reading
- Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
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