A live individual of Tylodina perversa in situ
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
(unranked): clade Heterobranchia
clade Euthyneura
clade Euopisthobranchia[1]
clade Umbraculida
Superfamily: Umbraculoidea
Dall, 1889 (1827)

See text

Umbraculoidea is a superfamily of unusual false limpets with a thin soft patelliform shell, marine gastropod molluscs in the clade Umbraculida, within the clade Euopisthobranchia.

There are two families in this superfamily, which is listed as the only superfamily in the clade Umbraculida within the informal group Opisthobranchia in the taxonomy of Bouchet & Rocroi (2005).[2]


A study by Grande et al., published in 2004, concluded that Umbraculoidea was a sister clade to the Cephalaspidea (Acteonoidea excluded).[3]

2005 taxonomy

Umbraculoidea contains two families:

2010 taxonomy

Jörger et al. (2010)[1] moved Umbraculoidea to the Euopisthobranchia.


  1. 1 2 Jörger K. M.; Stöger I.; Kano Y.; Fukuda H.; Knebelsberger T.; Schrödl M. (2010). "On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia". BMC Evolutionary Biology. 10: 323. doi:10.1186/1471-2148-10-323. PMC 3087543Freely accessible. PMID 20973994.
  2. Bouchet P.; Rocroi J.-P.; Frýda J.; Hausdorf B.; Ponder W.; Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology. Hackenheim, Germany: ConchBooks. 47 (1-2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  3. C. Grande; J. Templado; J.L. Ververa; R. Zardoya (2004). "Phylogenetic relationships among Opisthobranchia (Mollusca, Gastropoda) based on mitochondrial cox 1, tmV, and rmL genes". Molecular Phylogenetics and Evolution. 33 (2): 378–388. doi:10.1016/j.ympev.2004.06.008. PMID 15336672.
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