Pamphiliidae

Pamphiliidae

Acantholyda nemoralis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Suborder:	Symphyta
Superfamily:	Pamphilioidea
Family:	Pamphiliidae Cameron, 1890
Diversity
~200 species in 8 genera

Pamphiliidae (sometimes incorrectly spelled Pamphilidae) is a small wasp family within Symphyta, containing some 200 species from the temperate regions of North America and Eurasia. The larvae feed on plants (often conifers), using silk to either build webs or tents, or to roll leaves into tubes, in which they feed, thus earning them the common names leaf-rolling sawflies or web-spinning sawflies. They are distinguished from the closely related Megalodontesidae by their simple, filiform antennae.

Subfamilies and genera

The family is currently divided into three subfmailies based on phylogenetic analysis of both extant and extinct species.^[1]

Cephalciinae
- Acantholyda Costa, 1894
- Caenolyda Konow, 1897
- Cephalcia Panzer, 1805
- †Tapholyda Rasnitsyn, 1983
Juralydinae
- †Atocus Scudder, 1892
- †Juralyda Rasnitsyn, 1977
- Neurotoma Konow, 1897
- †Scabolyda Wang et al, 2014
Pamphiliinae
- Kelidoptera Konow, 1897
- Onycholyda Takeuchi, 1938
- Pamphilius Latreille, 1802
- Pseudocephaleia Zirngiebl, 1937
Incertae sedis
- †Ulteramus Archibald & Rasnitsyn, 2015

Extant Hymenopteran families

Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Subclass: Pterygota
Infraclass: Neoptera
Superorder: Endopterygota

S
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Xyeloidea	Xyelidae

Pamphilioidea	Megalodontesidae Pamphiliidae (web-spinning sawflies)

Tenthredinoidea	Argidae (argid sawflies) Blasticotomidae (fern sawflies) Cimbicidae (cimbicid sawflies) Diprionidae (conifer sawflies) Pergidae (pergid sawflies) Tenthredinidae (common sawflies)

U
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Cephoidea	Cephidae (stem sawflies)

Siricoidea	Anaxyelidae (cedar wood wasps) Siricidae (horntails)

Xiphydrioidea	Xiphydriidae (wood wasps)

Orussoidea	Orussidae (parasitic wood wasps)

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U
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P
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Stephanoidea

Stephanidae

Ichneumonoidea

Proctotrupomorpha

Platygastroidea	Platygastridae Scelionidae

Cynipoidea	Austrocynipidae Cynipidae (gall wasps) Figitidae Ibaliidae Liopteridae

Proctotrupoidea (s.str.)	Proctorenyxidae Roproniidae Heloridae Pelecinidae Peradeniidae Proctotrupidae Vanhorniidae

Diaprioidea	Austroniidae Diapriidae Maamingidae Monomachidae

Mymarommatoidea	Mymarommatidae

Chalcidoidea (chalcid wasps)	Agaonidae (fig wasps) Aphelinidae Chalcididae Encyrtidae Eucharitidae Eulophidae Eupelmidae Eurytomidae Leucospidae Mymaridae (fairyflies) Ormyridae Perilampidae Pteromalidae Rotoitidae Signiphoridae Tanaostigmatidae Tetracampidae Torymidae Trichogrammatidae

Ceraphronoidea

Megalyroidea

Megalyridae

Trigonaloidea

Trigonalidae

Evanioidea

A
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l
e
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Apoidea

Anthophila (bees)	Andrenidae (mason bees) Apidae (honey bees, bumblebees, cuckoo bees, carpenter bees, orchid bees, stingless bees) Colletidae (plasterer bees) Halictidae (sweat bees) Megachilidae (mason bees, leafcutter bees) Melittidae Stenotritidae

Spheciformes (sphecoid wasps)	Ampulicidae (cockroach wasps) Crabronidae (sand wasps, bee wolves) Heterogynaidae Sphecidae (thread-waisted wasps)

Chrysidoidea

Vespoidea

Italic are paraphyletic groups

Based on Malm and Nyman (2015)

References

↑ Wang, M.; Rasnitsyn, A.P.; Li, H.; Shih, C.; Sharkey, M.J.; Ren, D. (2015). "Phylogenetic analyses elucidate the inter‐relationships of Pamphilioidea (Hymenoptera, Symphyta)". Cladistics. 32 (3): 239–260. doi:10.1111/cla.12129.

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