Caryophyllidia are an anatomical feature of the external dorsal surface of dorid sea slugs. Caryophyllidia are sensory tubercles, surrounded by tiny needle-like structures called spicules, that are present on the outer mantle.[1][2][3]


  1. "". Archived from the original on 2011-08-27.
  2. "Marine species identification portal glossary".
  3. Camacho-García, Y.E.; Valdés, Á. (2003). "Caryophyllidia-bearing dorid nudibranchs (Mollusca, Nudibranchia, Doridacea) from Costa Rica" (PDF). California Academy of Sciences.
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