Bryan Clarke

Professor Bryan Campbell Clarke FRS,^[1] (24 June 1932 – 27 February 2014) was a British geneticist, latterly professor emeritus of genetics at the University of Nottingham. Clarke is particularly noted for his work on apostatic selection (which is a term he coined in 1962) and other forms of frequency-dependent selection, and work on polymorphism in snails, much of it done during the 1960s. Later, he studied molecular evolution. He made the case for natural selection as an important factor in the maintenance of molecular variation, and in driving evolutionary changes in molecules through time. In doing so, he questioned the over-riding importance of random genetic drift advocated by King, Jukes, and Kimura. With Professor James J Murray Jnr (University of Virginia), he carried out an extensive series of studies on speciation in land snails of the genus Partula inhabiting the volcanic islands of the Eastern Pacific. These studies helped illuminate the genetic changes that take place during the origin of species.

Academic career

Clarke was educated at Magdalen College Oxford, receiving a BA in 1956 and DPhil in 1961. He was appointed as Assistant Lecturer at Edinburgh University in 1959 and had been promoted to Reader by the time he left in 1971. In 1971 he became Foundation Professor at the new Department of Genetics at the University of Nottingham becoming Emeritus Professor in 1997. During this period he spent two spells (1971–76, 1981–93) as Head of Department.

Clarke mentored many scientists in evolutionary genetics, supervising more than thirty research students, many of which have gone on to successful research careers themselves. He was a co-founder of the Population Genetics Group ("PopGroup") a scientific meeting for evolutionary and population genetics held annually in the UK since the 1960s.^[2]

Clarke was co-founder and trustee of the Frozen Ark project to preserve the DNA and living cells of endangered species worldwide.

Clarke acted as managing editor of the journal Heredity from 1978 to 1985.^[3]

Awards and achievements

Clarke was elected a Fellow of the Royal Society in 1982, awarded the Linnean Medal for Zoology and elected a Foreign member of the American Philosophical Society in 2003. He was elected a Foreign Honorary Member of the American Academy of Arts and Sciences in 2004. He was one of thirteen recipients of the Darwin-Wallace Medal in 2008; this award is given every 50 years by the Linnean Society of London. He was awarded the Darwin Medal of the Royal Society in 2010 'for his original and influential contributions to our understanding of the genetic basis of evolution'.^[4]

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